古似鳥龍屬

古似鸟龙属
化石时期:白堊紀晚期
96 Ma
亞洲古似鳥龍的骨架模型,位於中國古動物博物館
科学分类 编辑
界: 动物界 Animalia
门: 脊索动物门 Chordata
纲: 蜥形纲 Sauropsida
总目: 恐龍總目 Dinosauria
目: 蜥臀目 Saurischia
亚目: 獸腳亞目 Theropoda
演化支 似鳥龍類 Ornithomimosauria
科: 似鸟龙科 Ornithomimidae
属: 古似鸟龙属 Archaeornithomimus
Russell英语Dale Russell, 1972
模式種
亞洲古似鳥龍
Archaeornithomimus asiaticus

(Gilmore, 1933[原本是亞洲似鳥龍])
其他
  • 比斯克提古似鳥龍 Archaeornithomimus bissektensis
    Nessov, 1995

古似鳥龍屬名Archaeornithomimus,意為「古老的鳥類模仿者」)是一似鳥龍下目獸腳亞目恐龍,生存於白堊紀晚期(約9600萬年前)的亞洲,化石來自二連組

發現及命名

1923年,由羅伊·查普曼·安德魯斯率領的美國自然史博物館團隊至內蒙古考察期間,彼得·凱森在三個採石場發現數個獸腳類遺骸,由多個個體組成,大部分關節分離,並缺乏頭骨和下頜。這些化石由查爾斯·惠特尼·吉爾摩於1933年命名並簡短敘述為似鳥龍屬的新種:亞洲似鳥龍(Ornithomimus asiaticus);種名意指來自亞洲。[1]1972年,戴爾·羅素英语Dale Russell將這個種建立成新的屬古似鳥龍,使亞洲古似鳥龍Archaeornithomimus asiaticus)成為其模式種。屬名由似鳥龍的屬名前面加上希臘文的ἀρχαῖος(archaios,「古老、原始」)組成,因為羅素認為發現古似鳥龍的地層屬於森諾曼階土侖階時期,約9500萬年前,使其成為當時已知最古老的似鳥龍科[2]最初吉爾摩並未編出一個正模標本;1990年,大衛·史密斯和彼得·加爾東在首次對化石的全面性敘述中,選擇了AMNH 6565標本(一隻腳)作為選定模式標本英语lectotype[3]

發現於馬里蘭州阿倫德爾組英语Arundel Formation的腳部骨頭曾於1888年由奧斯尼爾·查爾斯·馬許歸入異特龍Allosaurus medius)。[4]1911年,理察·斯旺·盧爾英语Richard Swann Lull將這個標本命名為傷龍的新種(Dryptosaurus grandis)。[5]1920年,吉爾摩將它們重新命名為似鳥龍的新種,但因為Ornithomimus grandis這個名稱已經存在,他將種名更名為Ornithomimus affinis[6]1972年,羅素將它們更名為古似鳥龍的第二個種(Archaeornithomimus affinis)。[2]但1990年史密斯和加爾東認為這些遺骸不屬於似鳥龍類,而是來自其他某些小型獸腳類。[3]

1995年,可能的第三個種:比斯克提古似鳥龍Archaeornithomimus bissektensis)由Lev A. Nesov根據發現於烏茲別克比斯克提組英语Bissekty Formation(土侖階-科尼亞克階)的模式標本N 479/12457:一個少年體的股骨蹠骨而命名。[7]儘管如此,其親緣關係普遍受到質疑或被忽視。[8][9][10]

敘述

與平均男性人類的體型比例圖
復原圖

古似鳥龍是種中型似鳥龍類,身長3.4公尺、體重介於45至91公斤。[11]來自其他似鳥龍類親緣物種的確切證據表明,古似鳥龍身上也覆有類似平胸鳥類的羽毛[12][13]以及角質喙狀嘴。[14][15]

後肢結構堅實。第三蹠骨近端未癒合,因此非併蹠骨型。[3]頸椎大幅氣腔化英语Skeletal pneumaticity,沿神經弓及椎體有非常複雜的內部空腔,代表具有頸部氣囊英语Air sac。前部胸椎和一些尾椎也有一定程度的氣腔化,而薦骨則未氣腔化。[9]2001年布魯斯·羅斯柴爾德和同事的研究試圖檢驗229件編入古似鳥龍的腳骨是否有出現疲勞性骨折的跡象,但結果是否定的。[16]

分類

羅素將古似鳥龍編入似鳥龍科[2]近期的系統發生學分析要麼支持這種分類要麼將本屬排除於似鳥龍科之外、位於似鳥龍下目的基礎位置。以下演化樹取自2019年似鳥身女妖龍敘述論文中的廣泛虛骨龍類系統發育分析,古似鳥龍在這裡被分類至似金翅鳥龍科,與阿肯色龍近緣。[10]

內蒙古博物館的骨架
似鳥龍下目 Ornithomimosauria

鶴形龍 Hexing

恐手龍 Deinocheirus

神州龍 Shenzhousaurus

昂雅克夏朗德的分類單元 Angeac taxon

內德科爾伯特龍 Nedcolbertia

似鳥龍科 Ornithomimidae

似象鳥龍 Aepyornithomimus

中國似鳥龍 Sinornithomimus

似鳥龍 Ornithomimus

似金翅鳥龍科 Garudimimidae

阿肯色龍 Arkansaurus

古似鳥龍 Archaeornithomimus

GIN 960910KD標本

北山龍 Beishanlong

似金翅鳥龍 Garudimimus

似鳥身女妖龍 Harpymimus

蒙古似雞龍 "Gallimimus mongoliensis"

似多多鸟龙 Tototlmimus

MNAP 11762A標本

前似鴕龍 Rativates

坐似鳥龍 Ornithomimus sedens

似鵝龍 Anserimimus

秋扒龍 Qiupalong

似馳龍 Dromiceiomimus

似鴕龍 Struthiomimus

似提姆龍 Timimus

似雞龍 Gallimimus

古生態學

兩隻巨盜龍保護牠們的巢穴免受獨龍古似鳥龍騷擾

古似鳥龍遺骸發現於二連組,年代可追溯至白堊紀晚期森諾曼階,約9600萬年前。[17]地層所呈現的環境是充滿辮狀河流曲流[18][19]、與海洋接壤[20]氾濫平原,並如古土壤英语Paleosol發展所示支持著廣泛的植被以及大量的植食性恐龍(如鴨嘴龍科)。[18]

如同其他似鳥龍類,古似鳥龍可能是雜食性動物,具有角質嘴喙,從小哺乳類、植物、水果、蛋、到甚至是其他恐龍幼體都吃。[14][15]

與古似鳥龍共棲的恐龍包括:其他獸腳類如暴龍超科獨龍鐮刀龍科二連龍內蒙古龍偷蛋龍科巨盜龍;植食性恐龍以鴨嘴龍類巴克龍計氏龍蜥腳類蘇尼特龍為代表。[21]

參見

似鳥龍類研究歷史英语Timeline of ornithomimosaur research

參考資料

  1. ^ Gilmore, C. W. On the dinosaurian fauna of the Iren Dabasu Formation. Bulletin of the American Museum of Natural History. 1933, 67 (2): 23–78. hdl:2246/355. 
  2. ^ 2.0 2.1 2.2 Russell, D. A. Ostrich dinosaurs from the Late Cretaceous of Western Canada. Canadian Journal of Earth Sciences. 1972, 9 (4): 375–402 [2020-12-30]. doi:10.1139/e72-031. (原始内容存档于2021-01-17). 
  3. ^ 3.0 3.1 3.2 Smith, D.; Galton, P. Osteology of Archaeornithomimus asiaticus (Upper Cretaceous, Iren Dabasu Formation, People's Republic of China). Journal of Vertebrate Paleontology. 1990, 10 (2): 255–265. doi:10.1080/02724634.1990.10011811. 
  4. ^ Marsh, O. C. Notice of a new genus of Sauropoda and other new dinosaurs from the Potomac Formation. American Journal of Science. Series 3. 1888, 35 (205): 89–94 [2020-12-30]. Bibcode:1888AmJS...35...89M. doi:10.2475/ajs.s3-35.205.89. (原始内容存档于2021-01-17). 
  5. ^ Lull, R. S. The Reptilia of the Arundel Formation. Maryland Geological Survey: Lower Cretaceous. 1911: 173–178. 
  6. ^ Gilmore, C. W. Osteology of the carnivorous dinosauria in the United States National Museum, with special reference to the genera Antrodemus (Allosaurus) and Ceratosaurus. Bulletin of the United States National Museum. 1920, 110 (110): i–159. OCLC 1836981. doi:10.5479/si.03629236.110.i. hdl:10088/10107. 
  7. ^ Nesov, L. A. Dinozavri severnoi Yevrazii: Novye dannye o sostave kompleksov, ekologii i paleobiogeografii. Institute for Scientific Research on the Earth's Crust (St Petersburg State University). 1995: 1–156. 
  8. ^ Lee, Y.-N.; Barsbold, R.; Currie, P. J.; Kobayashi, Y.; Lee, H.-J.; Godefroit, P.; Escuillié, F.; Chinzorig, T. Resolving the long-standing enigmas of a giant ornithomimosaur Deinocheirus mirificus. Nature. 2014, 515 (7526): 257–260. Bibcode:2014Natur.515..257L. PMID 25337880. doi:10.1038/nature13874. 
  9. ^ 9.0 9.1 Watanabe, A.; Gold, M. E. L. G.; Brusatte, S. L.; Benson, R. B. J.; Choiniere, J.; Davidson, A.; Norell, M. A. Vertebral Pneumaticity in the Ornithomimosaur Archaeornithomimus (Dinosauria: Theropoda) Revealed by Computed Tomography Imaging and Reappraisal of Axial Pneumaticity in Ornithomimosauria. PLOS ONE. 2015, 10 (12): e0145168. PMC 4684312可免费查阅. PMID 26682888. doi:10.1371/journal.pone.0145168. 
  10. ^ 10.0 10.1 Hartman, S.; Mortimer, M.; Wahl, W. R.; Lomax, D. R.; Lippincott, J.; Lovelace, D. M. A new paravian dinosaur from the Late Jurassic of North America supports a late acquisition of avian flight. PeerJ. 2019, 7: e7247. PMC 6626525可免费查阅. PMID 31333906. doi:10.7717/peerj.7247. 
  11. ^ Holtz, T. R.; Rey, L.V. Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages. Random House. 2007. ISBN 9780375824197.  Genus List for Holtz 2012页面存档备份,存于互联网档案馆Weight Information页面存档备份,存于互联网档案馆
  12. ^ Zelenitsky, D. K.; Therrien, F.; Erickson, G. M.; DeBuhr, C. L.; Kobayashi, Y.; Eberth, D. A.; Hadfield, F. Feathered Non-Avian Dinosaurs from North America Provide Insight into Wing Origins. Science. 2012, 338 (6106): 510–514. PMID 23112330. doi:10.1126/science.1225376. 
  13. ^ Van der Reest, A. J.; Wolfe, A. P.; Currie, P. J. A densely feathered ornithomimid (Dinosauria: Theropoda) from the Upper Cretaceous Dinosaur Park Formation, Alberta, Canada. Cretaceous Research. 2016, 58: 108–117. doi:10.1016/j.cretres.2015.10.004. 
  14. ^ 14.0 14.1 Norell, M. A.; Makovicky, P. J.; Currie, P. J. The beak of ostrich dinosaurs. Nature. 2001, 412 (6850): 873–874. PMID 11528466. doi:10.1038/35091139. 
  15. ^ 15.0 15.1 Barrett, P. M. The diet of ostrich dinosaurs (Theropoda: Ornithomimosauria). Palaeontology. 2005, 48 (2): 347–358. doi:10.1111/j.1475-4983.2005.00448.x. 
  16. ^ Rothschild, B.; Tanke, D. H.; Ford, T. L. Theropod stress fractures and tendon avulsions as a clue to activity. Tanke, D. H.; Carpenter, K.; Skrepnick, M. W. (编). Mesozoic Vertebrate Life. Indiana University Press. 2001: 331–336. ISBN 9780253339072. 
  17. ^ Guo, Z. X.; Shi, Y. P.; Yang, Y. T.; Jiang, S. Q.; Li, L. B.; Zhao, Z. G. Inversion of the Erlian Basin (NE China) in the early Late Cretaceous: Implications for the collision of the Okhotomorsk Block with East Asia (PDF). Journal of Asian Earth Sciences. 2018, 154: 49–66 [2020-12-30]. doi:10.1016/j.jseaes.2017.12.007. (原始内容 (PDF)存档于2020-09-19). 
  18. ^ 18.0 18.1 Van Itterbeeck, J.; Horne, D. J.; Bultynck, P.; Vandenberghe, N. Stratigraphy and palaeoenvironment of the dinosaur-bearing Upper Cretaceous Iren Dabasu Formation, Inner Mongolia, People's Republic of China. Cretaceous Research. 2005, 26 (4): 699–725 [2020-12-30]. doi:10.1016/j.cretres.2005.03.004. (原始内容存档于2021-12-05). 
  19. ^ Averianov, A.; Sues, H. Correlation of Late Cretaceous continental vertebrate assemblages in Middle and Central Asia (PDF). Journal of Stratigraphy. 2012, 36 (2): 462–485 [2020-12-30]. (原始内容 (PDF)存档于2019-03-07). 
  20. ^ Funston, G. F.; Currie, P. J.; Ryan, M. J.; Dong, Z.-M. Birdlike growth and mixed-age flocks in avimimids (Theropoda, Oviraptorosauria). Scientific Reports. 2019, 9 (18816): 18816. PMC 6906459可免费查阅. PMID 31827127. doi:10.1038/s41598-019-55038-5. 
  21. ^ Xing, H.; He, Y.; Li, L.; Xi, D. A review on the study of the stratigraphy, sedimentology, and paleontology of the Iren Dabasu Formation, Inner Mongolia. Wei, D. (编). Proceedings of the Thirteenth Annual Meeting of the Chinese Society of Vertebrate Paleontology. Beijing: China Ocean Press. 2012: 1–44 (中文). 

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Dutch politician and diplomat In this Dutch name, the surname is De Vries, not Vries. Gijs de VriesMember of the European Court of AuditorsIn office1 January 2011 – 1 January 2014Member of the Court of AuditIn office1 January 2008 – 1 January 2011European Union Counter-terrorism CoordinatorIn office25 March 2004 – 1 March 2007Secretary-GeneralJavier SolanaPreceded byOffice establishedSucceeded byGilles de KerchoveMember of the House of RepresentativesIn office...

 

UK phone retailer This article needs additional citations for verification. Please help improve this article by adding citations to reliable sources. Unsourced material may be challenged and removed.Find sources: Dial-a-Phone – news · newspapers · books · scholar · JSTOR (November 2014) (Learn how and when to remove this message) DialaPhoneIndustryTelecommunicationsFounded1995FounderJonathan Beck and Richard FrankDefunct15 September 2014FateDefunct (ba...