Royal Society (
1987–)
[1]Det franske vitenskapsakademiet (
2000–)
[2]Det pavelige vitenskapsakademi (
1978–)
[3]National Academy of Sciences (
1974–)
[4]American Academy of Arts and Sciences (
1974–)
[5]American Association for the Advancement of Science (
1980–)
[3]European Molecular Biology Organization (
1983–)
[6]Academia Europaea (
1999–) (tilknytning: AE section Cell and developmental biology)
[7]American Medical Writers Association (
1985)
[4]National Academy of Medicine (
1988)
[4]The Japanese Biochemical Society (
1991)
[4]American Academy of Microbiology (
1992)
[4]