Initially the squadron was equipped with Hawker Hurricane I fighters.[1] The squadron was operational in only a month and as part of 12 Group[2] took part in the Battle of Britain as part of the DuxfordBig Wing. The squadron claimed 37½ victories in the battle.
From 1941 the squadron began offensive operations flying sweeps over the English Channel and providing bomber escorts. In 1941 Hess was transferred to be a Czechoslovak liaison officer at Fighter Command, and on 28 February Sqn Ldr František Weber [cs] succeeded him in command of 311 Squadron.[4]
On 14 December 1941 the squadron moved to RAF Perranporth[7] in Cornwall for defensive operations. On 7 April 1942 Sqn Ldr František Doležal succeeded Weber as squadron commander. On 7 May the squadron moved to RAF Exeter.[7]
On 15 January 1943 Doležal was succeeded by Sqn Ldr Emil Foit.[4] On 26 June 1943 the squadron moved to RAF Castletown[7] in Caithness, Scotland for another three-month rest period. From July to September it operated the Spitfire VI.[6] On 18 September 1943 the squadron moved to RAF Ibsley[7] in Hampshire.
On 13 January 1944 Sqn Ldr Hugo Hrbáček succeeded Foit as squadron commander.[5] On 20 February the squadron moved to RAF Mendlesham in Suffolk, on 29 March it moved again to RAF Rochford in Essex, and on 3 April it moved again to RAF Appledram in West Sussex.[7] On 21 May Sqn Ldr Václav Raba succeeded Hrbáček as squadron commander.[5]
In 1944 the squadron was re-equipped with the Spitfire IX and became a fighter-bomber unit with 134 Wing, flying ground attack duties during the Normandy landings. From 22 June until 4 July the squadron was based at RAF Tangmere in West Sussex, and from 4 to 11 July it was based at RAF Lympne in Kent.[7]
On 11 July 1944 the squadron moved to RAF Digby in Lincolnshire, and on 28 August it moved again to RAF North Weald in Essex.[8] On 15 September Sqn Ldr Jiří Hartman succeeded Raba as squadron commander.[5] The squadron then spent the rest of the war flying armed reconnaissance missions along the Dutch and Belgian coasts. On 29 December 1944 it moved to RAF Bradwell Bay in Essex, and on 27 February 1945 the squadron moved to RAF Manston in Kent.[8]
The squadron's final score was 52½ claims including four V-1 flying bombs shot down. Three of those flying bombs were shot down on 8 July by the same fighter ace, Flt LtOtto Smik DFC, in one sortie, within 32 minutes of each other.[9]
On 7 August 1945 the squadron moved to Hildesheim, and on 31 August it moved again to Ruzyně Airport in Prague.[8] It became a squadron of the new Czechoslovak Air Force,[6] and on 15 February 1946 was officially disbanded as an RAF squadron.[1]
Supermarine Spitfire Vc 'AR501' (civil registration G-AWII) built in 1942, remains airworthy, and is maintained & operated by The Shuttleworth Collection in Bedfordshire, England. AR501 was one of a production batch of 300 Spitfires ordered under Contract No. 1305/40 of August 1940 from Westland Aircraft, Yeovil.
Following restoration in 2018 by The Shuttleworth Collection it is presented in the squadron colours of No. 312 (Czech) Squadron, which the aircraft moved to on October 10th 1943 from No. 310 (Czech) Squadron. Shuttleworth periodically switch between elliptical and clipped wing configurations.
Halley, James J (1988). The Squadrons of the Royal Air Force & Commonwealth, 1918–1988. Tonbridge: Air-Britain (Historians) Ltd. ISBN0-85130-164-9.
Hurt, Zdeněk (2004). In Focus: Czechs in the RAF. Walton-on-Thames: Red Kite. ISBN0-9538061-9-7.
Jackson, Robert (2003). Spitfire The History of Britain's Most Famous World War II Fighter. Bath: Parragon. p. 89. ISBN0-75258-770-6.
Jefford, Wg Cdr CG (2001) [1998]. RAF Squadrons, A Comprehensive Record of the Movement and Equipment of all RAF Squadrons and their Antecedents since 1912 (second ed.). Shrewsbury: Airlife Publishing. ISBN978-1-84037-141-3.
Lewis, Peter (1968) [1959]. Squadron Histories, RFC, RNAS and RAF, Since 1912. London: Putnam. p. 96. SBN 370-00022-6.
Rawlings, John DR (1976) [1969]. Fighter Squadrons of the RAF and their Aircraft (new ed.). London: Macdonald and Jane's (Publishers) Ltd. ISBN0-354-01028-X.
van Eyck, Manuel F (1993). Zemřeli jsme pro Anglii (in Czech). Translated by František Fajtl. prologue by František Fajtl. Prague: Naše vojsko. ISBN80-206-0321-2.