Scutelleridae

Jewel bugs
The shield bug, Scutiphora pedicellata, on a leaf.
Scutiphora pedicellata, a jewel bug from Australia
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Hemiptera
Suborder: Heteroptera
Infraorder: Pentatomomorpha
Superfamily: Pentatomoidea
Family: Scutelleridae
Leach, 1815
Subfamilies

Scutelleridae is a family of true bugs. They are commonly known as jewel (stink) bugs or metallic shield bugs due to their often brilliant coloration. With the name based on the Asian genus Scutellera, they are also known as shield-backed bugs due to the enlargement of the thoracic scutellum into a continuous shield over the abdomen and wings.[1] This latter characteristic distinguishes them from most other families within Heteroptera, and may lead to misidentification as a beetle rather than a bug. These insects feed on plant juices from a variety of different species, including some commercial crops. Closely related to stink bugs, they may also produce an offensive odour when disturbed. There are around 450 species worldwide.[2]

Description

Jewel bugs are small to medium-sized oval-shaped bugs with a body length averaging at 5 to 20 mm (0.20 to 0.79 in).[3] They can easily be distinguished from stink bugs (Pentatomidae) because the shield-like enlarged last section of their thorax (known as the scutellum, Latin for "little shield") completely covers the abdomen and the wings.[4]

Despite their resemblance to beetles, jewel bugs are hemipterans or true bugs.[5] The scutellum is an extension of the thorax, unlike the elytra of beetles which are hardened forewings. As such, jewel bugs have four membranous wings underneath the scutellum in contrast to two in beetles.[6] The scutellum in jewel bugs also does not have a division in the middle and thus does not 'split open' when they take flight like in beetles.[7]

The heads of jewel bugs are triangular and the antennae have three to five segments.[8] Like all heteropterans, jewel bugs are characterized by a segmented beak-like mouthpart (known as the rostrum).[2] During feeding, jewel bugs inject proteolytic enzymes in their saliva into plants, digesting plant matter into a liquid form which they then suck up.[9] The tarsus has three segments (tarsomeres).[6]

Colours

Though some species are quite drab,[10] the most conspicuous jewel bugs are often brilliantly colored, exhibiting a wide range of iridescent metallic hues that change with the view angle.[6] The colors are the result of a combination of factors. Some species like Chrysocoris stockerus and Scutellera nobilis display colors from multiple thin layers of pigmented chitin. The colors often change or become duller when the specimens are dried, due to the topmost chitinous layer becoming opaque and obscuring the colors of the bottom layer. The colors can be restored by moistening the surfaces with water.[11]

Iridescence (or goniochromism) in jewel bugs like Poecilocoris lewisi are the result of structural coloration. Instead of pigments, the colors are caused by the interference, diffraction, or scattering of light by numerous tiny structures.[citation needed]

In Poecilocoris lewisi, multiple tiny conical protuberances around 900 nm in height and averaging at a diameter of 360 nm are scattered on the epicuticle. These structures affect light passing through them, producing their oily-looking blue sheen (known as the Tyndall effect or Mie scattering).[11]

In other species like the African shield bug (Calidea panaethiopica), the dorsal cuticle is dotted with tiny regularly spaced hemispherical cavities. The depressions act like Bragg mirrors. When light hits the pitted surface, it gives off multiple reflections resulting in the distinctive two tone yellow-blue iridescence.[12]

The colors and patterns on jewel bugs can vary significantly between instars and even within adults of a species.[13]

Jewel bugs are also known to mimic the colors, patterns, and shape of other organisms for defensive purposes. An example is the yellow-spotted black Steganocerus multipunctatus which exhibits Müllerian mimicry with the tortoise beetle Chiridopsis suffriani.[14]

Ecology and life cycle

All jewel bugs feed on plants (phytophagous). The eggs are laid in compact clusters.[15] They may be round or barrel-shaped with a lid or a cap at the top (known as the operculum). They also contain a ring of small protuberances near the cap called micropylar processes. They permit the passage of sperm into the egg for fertilization and enable gaseous exchange from within the egg and the outside world for the embryos. The eggs are white or cream colored when freshly laid but can change color as the embryo matures.[16] When hatching, the prolarva (the advanced embryo) exit the egg by opening the lid through peristaltic movements and with the help of a T-shaped internal structure in the egg (known as the egg burster).[16][17]

Like all hemipterans, jewel bugs undergo incomplete metamorphosis (hemimetaboly) and do not possess larval and pupal stages. Instead the adults develop from several stages (instars) of nymphs (usually five) through successive moltings (ecdysis). Nymphs resemble the adults except for size and the absence of wings. They can be of different coloration or patterns from adults.[15]

Some species are known to exhibit parental care of eggs and nymphs. Notable examples of which are Cantao parentum,[18] Pachycoris klugii,[19] Pachycoris stalii,[20] Pachycoris torridus, and Tectocoris diophthalmus.[13][21]

Reproduction

Chemical secretions from dorsal abdominal or sternal exocrine glands are used to attract mates by certain species of jewel bugs.[22] In certain genera (like Tectocoris, Psacasta, Odontoscelis, and Irochrotus), males possess special unicellular glands in the abdomen known as the androconia (singular: androconium). They release sex pheromones when ruptured.[6]

Females possess a spermatheca, an ectodermal gland which opens into the oviduct. These serve as storage for sperm deposited by males. It contains glands which can nourish the spermatozoa until they can be released to fertilize eggs.[17]

Male jewel bugs of the genus Hotea possess an unusually large, spiky, and heavily sclerotized genitalia. They are used in a mating practice known as traumatic insemination, a result of evolutionary sexual conflict. Male Hotea bugs tear through the female reproductive ducts to deposit sperm, inflicting substantial damage to the female in the process.[23]

Defenses

Like stink bugs, a vast majority of jewel bugs, both adults and nymphs, are also capable of releasing pungent defensive chemicals from glands located on the sides of the thorax.[2][7] Typical compounds exuded by jewel bugs include alcohols, aldehydes, and esters.[24]

Nymphs and adults often exhibit clustering behavior, being found in large numbers close to each other. This behavior is thought to have an evolutionary advantage. The more individuals present in an area, the stronger the odor of the chemicals released when the bugs are threatened.[18] If this fails, stink bugs will react to threat by flying away or dropping to the ground.[2]

Classification and evolution

Scutellerids were first described by the English zoologist William Elford Leach in 1815.[25][26] It belongs to the order Hemiptera (true bugs), under the suborder Heteroptera and infraorder Pentatomomorpha. They are classified under the superfamily Pentatomoidea. They were formerly classified as a subfamily of Pentatomidae by George Willis Kirkaldy in 1909.[27] The earliest attempt to restore them to family status was in 1917 by Edward Payson Van Duzee.[28] Most authorities today regard it as a valid family group.[27] In phylogenetic studies in 2008 by Grazia et al., Scutelleridae was shown to be consistently monophyletic, basal to Acanthosomatidae, and distal to Plataspididae and Parastrachiidae.[28][29] Below is the morphological unweighted tree of the superfamily Pentatomoidea after Grazia et al. (2008).[29]

      ←      
             

Urostylididae

             
             

Saileriolidae

             
             

Acanthosomatidae

             
             
             

Tessaratomidae

             

Dinidoridae

  Cydnidae sensu lato  
             

Cydnidae

             

Thaumastellidae

             

Parastrachiinae

             
             

Thyreocoridae

             

Lestoniidae

             

Phloeidae

             
             

Scutelleridae

             

Plataspididae

             

Pentatomidae

             

Canopidae

             

Megarididae

Subfamilies and genera

The family is composed of about 81 genera and around 450 species worldwide. While the tribal and subfamilial classifications remain unclear,[30] they are divided into eight subfamilies sensu lato: Elvisurinae, Eurygastrinae, Hoteinae (sometimes classified under Pachycorinae), Odontoscelinae, Odontotarsinae, Pachycorinae, Scutellerinae, and Tectocorinae.[31][32]

Elvisurinae

Auth.: Stål, 1872. Widespread distribution, with the type genus Elvisura found in southern Africa; only one species of Elvisurinae, Solenosthedium bilunatum, is found in Europe.[33]

  1. Austrotichus Gross, 1975 - monotypic A. rugosus Gross, 1975
  2. Coleotichus White, 1839
  3. Elvisura Spinola, 1837
  4. Solenosthedium Spinola, 1837
  5. Solenotichus Martin, 1897

Eurygastrinae

Auth.: Amyot & Audinet-Serville, 1843

Tribe Eurygastrini Amyot & Audinet-Serville, 1843
  1. Eurygaster Laporte, 1833
  2. Polyphyma Jakovlev, 1877
Tribe Psacastini Mulsant & Rey, 1865
  1. Ceratocranum Reuter, 1890
  2. Periphima Jakovlev, 1889
  3. Periphymopsis Schouteden, 1904
  4. Promecocoris Puton, 1886
  5. Psacasta Germar, 1839
  6. Xerobia Stål, 1873

Hoteinae

Auth.: Carapezza, 2008

  1. Deroplax Mayr, 1864
  2. Ellipsocoris Mayr, 1864
  3. Hotea Amyot & Serville, 1843

Odontoscelinae

Auth.: Amyot & Audinet-Serville, 1843; mostly European

  1. Holonotellus Horváth, 1895
  2. Irochrotus Amyot & Serville, 1843
  3. Odontoscelis Laporte, 1833

Odontotarsinae

Auth.: Mulsant & Rey, 1865

Tribe Odontotarsini Mulsant & Rey, 1865
  1. Ahmadocoris Carapezza, 2009
  2. Alphocoris Germar, 1839
  3. Melanodema Jakovlev, 1880
  4. Odontotarsiellus Hoberlandt, 1955
  5. Odontotarsus Laporte, 1833
  6. Urothyreus Horváth, 1921
Tribe Phimoderini Fuente, 1974
  1. Euptychodera Bergroth, 1908
  2. Fokkeria Schouteden, 1904
  3. Morbora Distant, 1899
  4. Phimodera Germar, 1839
  5. Vanduzeeina Schouteden, 1904

Pachycorinae

Auth.: Amyot & Audinet-Serville, 1843

  1. Acantholomidea Sailer, 1945
  2. Agonosoma Laporte, 1833
  3. Ascanius (bug) Stål, 1868
  4. Brailovskylus Eger, 2017
  5. Camirus Stål, 1862
  6. Chelycoris Bergroth, 1891
  7. Chelyschema Bergroth, 1891
  8. Coptochilus Amyot & Serville, 1843
  9. Crathis (bug) Stål, 1861
  10. Diolcus Mayr, 1864
  11. Dystus Stål, 1862
  12. Ephynes Stål, 1868
  13. Galeacius Distant, 1889
  14. Homaemus Dallas, 1851
  15. Lobothyreus Mayr, 1864
  16. Misippus Stål, 1868
  17. Nesogenes Horváth, 1921
  18. Orsilochides Kirkaldy, 1909
  19. Pachycoris Burmeister, 1835
  20. Polytes Stål, 1868
  21. Sphyrocoris Mayr, 1864
  22. Stethaulax Bergroth, 1891
  23. Symphylus Dallas, 1851
  24. Testrina Walker, 1867
  25. Tetyra Fabricius, 1803
  26. Tiridates (bug) Stål, 1868

Auth.: Leach, 1815. Selected genera:

Tectocorinae McDonald & Cassis, 1984 (monotypic)

Economic significance

Left: Eurygaster testudinaria, a sunn pest from Germany. Center: Odontotarsus grammicus, another sunn pest from Spain. Right: An adult and nymph cotton harlequin bug (Tectocoris diophthalmus).

Though most jewel bugs do little harm to crop plants,[34] a few members of Scutelleridae are considered major agricultural pests. Together with some species of stink bugs, they are collectively known as sunn pests (also spelled as senn, soun, or shüne pests) or wheat bugs.[30] The most economically important species of which are members of the genus Eurygaster.

Eurygaster integriceps, in particular, is a very destructive pest of cereal crops in North Africa, the Balkans, and western and central Asia. Other scutellerids known under the name 'sunn pest' include members of the genus Odontotarsus, among others.[35] Methods of control for sunn pests have included biological pest control, using wasps of the family Scelionidae from the genera Trissolcus and Ooencyrtus.[citation needed]

The cotton harlequin bug (Tectocoris diophthalmus) is also an important pest of cotton crops and Hibiscus.[34][36]

Conservation

Biological methods of pest control have sometimes backfired. A parasitoid fly which preys on hemipterans, Trichopoda pennipes was introduced to Hawaii to control the invasive species Nezara viridula, the southern green stink bug. The fly now threatens native species of bugs in Hawaii as well, particularly the Koa bug (Coleotichus blackburniae, a jewel bug species notable for not possessing stink glands) which has now become rare.[37]

See also

References

  1. ^ "Scutelleridae - Jewel Bugs". Commonwealth Scientific and Industrial Research Organisation (CSIRO) Entomology. Retrieved April 27, 2011.
  2. ^ a b c d John L. Capinera (2008). Encyclopedia of entomology. Springer. pp. 608–609. ISBN 978-1-4020-6242-1.
  3. ^ Robert G. Foottit & Peter Holdridge Adler (2009). Insect biodiversity: science and society. John Wiley and Sons. p. 247. ISBN 978-1-4051-5142-9.
  4. ^ Mike Picker; Charles Griffiths & Alan Weaving (2004). Field guide to insects of South Africa. Field Guide Series. Struik. p. 132. ISBN 978-1-77007-061-5.[permanent dead link]
  5. ^ "Scutelleridae". National Center for Biotechnology Information (NCBI).
  6. ^ a b c d Randall T. Schuh & James Alexander Slater (1995). True bugs of the world (Hemiptera:Heteroptera): classification and natural history. Cornell University Press. ISBN 978-0-8014-2066-5.
  7. ^ a b "Family Scutelleridae - Shield-backed Bugs". BugGuide. Retrieved April 27, 2011.
  8. ^ "Jewel bugs, Shield Backed Bugs -Family Scutelleridae". Brisbane Insects and Spiders Home Page. Retrieved April 27, 2011.
  9. ^ E. Grünbacher & B. Kromp (2008). "Investigations on the occurrence of wheat bugs (Scutelleridae, Pentatomidae; Heteroptera) in organic farming of Eastern Austria" (PDF). 1st Scientific Conference within the framework of the 8th European Summer Academy on Organic Farming. Retrieved April 27, 2011. {{cite journal}}: Cite journal requires |journal= (help)
  10. ^ Scott Fabricant, Mariella Herberstein & Darrell Kemp. "Mechanisms and Functions of Colour in the Hibiscus Harlequin Bug" (PDF). The Australasian Society for the Study of Animal Behaviour. Retrieved April 28, 2011. {{cite journal}}: Cite journal requires |journal= (help)
  11. ^ a b Shūichi Kinoshita (2008). Structural colors in the realm of nature. World Scientific. p. 155. ISBN 978-981-270-783-3.
  12. ^ Jean-Pol Vigneron, Moussa Ouedraogo, Jean-François Colomer, and Marie Rassart (200). "Spectral sideband produced by a hemispherical concave multilayer on the African shield-bug Calidea panaethiopica (Scutelleridae)". Physical Review E. 79 (2). American Physical Society: 021907. doi:10.1103/PhysRevE.79.021907. PMID 19391778.{{cite journal}}: CS1 maint: multiple names: authors list (link)
  13. ^ a b Luis Cervantes Peredo (2002). "Description, Biology, and Maternal Care of Pachycoris klugii (Heteroptera: Scutelleridae)" (PDF). Florida Entomologist. 85 (3). Instituto de Ecologia, Florida Center for Library Automation: 464–473. doi:10.1653/0015-4040(2002)085[0464:dbamco]2.0.co;2. Retrieved April 27, 2011.
  14. ^ Hugh D.C. Heron (2009). "Polymorphism in four tortoise beetles from Queensburgh, South Africa (Chrysomelidae: Cassidinae)" (PDF). Genus. 22 (1). University of Wrocław: 133–149. Retrieved April 28, 2011.
  15. ^ a b Maurice Burton & Robert Burton (1970). The international wildlife encyclopedia. Vol. 1. Marshall Cavendish. pp. 2344–2345. ISBN 978-0-7614-7266-7.
  16. ^ a b Selami Candan, Zekiye Suludere & Dilek Durak (2005). "Ultrastructure of the Eggs Chorion of Ceraleptus obtusus (Brulle, 1839)(Heteroptera: Coreidae)". Ohio Journal of Science. 105 (5). The Knowledge Bank, Ohio State University: 138–141. hdl:1811/31918.
  17. ^ a b Selami Candan, Zekiye Suludere & Mahmut Erbey (2008). "Morphology of eggs and spermatheca of Odontotarsus purpureolineatus (Heteroptera, Scutelleridae)". Biologia, Bratislava. 62 (6). Springer: 763–769. doi:10.2478/s11756-007-0137-x. ISSN 0006-3088.
  18. ^ a b "Scutelleridae: Shield-backed Bugs". Save Our Waterways Now. Retrieved April 28, 2011.
  19. ^ James T. Costa (2006). The other insect societies. Harvard University Press. p. 281. ISBN 978-0-674-02163-1.
  20. ^ Livy Williams III, Maria C Coscarón, Pablo M Dellapé, and Timberley M Roane (2005). "The shield-backed bug, Pachycoris stallii: Description of immature stages, effect of maternal care on nymphs, and notes on life history". J Insect Sci. 5 (29). University of Wisconsin Libraries: 29. doi:10.1093/jis/5.1.29. PMC 1615236. PMID 17119611.{{cite journal}}: CS1 maint: multiple names: authors list (link)
  21. ^ David A. Rider. "Pentatomoid Species Known to Exhibit Parental Care". Department of Entomology, North Dakota State University. Retrieved April 27, 2011.
  22. ^ Ring T. Cardé & William J. Bell (1995). Chemical ecology of insects 2. Springer. p. 336. ISBN 978-0-412-03961-4.
  23. ^ Göran Arnqvist & Locke Rowe (2005). Sexual conflict. Princeton University Press. p. 129. ISBN 978-0-691-12218-2.
  24. ^ Stefan Schulz (2005). The chemistry of pheromones and other semiochemicals. Vol. 2. Springer. p. 37. ISBN 978-3-540-21308-6.
  25. ^ "Scutelleridae". Integrated Taxonomic Information System.
  26. ^ "Scutelleridae Leach, 1815". The Encyclopedia of Life.
  27. ^ a b G. Cassis, Australia. Bureau of Flora and Fauna, & Gordon F. Gross (2002). Zoological catalogue of Australia: Hemiptera: Heteroptera (Pentatomomorpha). CSIRO Publishing. p. 353. ISBN 978-0-643-06875-9.{{cite book}}: CS1 maint: multiple names: authors list (link)
  28. ^ a b Jocelia Grazia; Randall T. Schuhb & Ward C. Wheeler (2008). "Phylogenetic relationships of family groups in Pentatomoidea based on morphology and DNA sequences (Insecta: Heteroptera)" (PDF). Cladistics. 24 (6). Wiley-Blackwell: 932–976. doi:10.1111/j.1096-0031.2008.00224.x. PMID 34892882. S2CID 41951432. Retrieved April 27, 2011.
  29. ^ a b Dimitri Forero (March 13, 2009). "Pentatomoidea". Tree of Life Web Project. Retrieved April 28, 2011.
  30. ^ a b Carl W . Schaefer; John D . Lattin & M . Javahery (2001). "Shield Bugs (Scutelleridae)". In Carl W . Schaefer & Antonio Ricardo Panizzi (eds.). Heteroptera of Economic Importance. CRC Press. pp. 475–503. doi:10.1201/9781420041859.ch14 (inactive 2024-11-12). ISBN 978-1-4200-4185-9.{{cite book}}: CS1 maint: DOI inactive as of November 2024 (link)
  31. ^ Biolib: family of shield-backed bugs, Scutelleridae Leach, 1815 (retrieved 10 February 2024)
  32. ^ David A. Rider. "Classification". Department of Entomology, North Dakota State University. Retrieved April 27, 2011.
  33. ^ Škorput, Jadranka; Martinović, Matea; van der Heyden, Torsten; Skejo, Josip (March 2020). "Solenosthedium bilunatum (Heteroptera: Scutelleridae) at the Adriatic Coast of Croatia". ResearchGate. Entomologie heute 31 (2019): 25-29. Retrieved 4 February 2024.
  34. ^ a b Murray Fletcher (2007). "Plant bugs" (PDF). Primefact. 508. NSW Department of Primary Industries: 1–4. Retrieved April 27, 2011.
  35. ^ Randall T. Schuh & James Alexander Slater (1995). True bugs of the world (Hemiptera:Heteroptera): classification and natural history. Cornell University Press. p. 36. ISBN 978-0-8014-2066-5.
  36. ^ Peter T. Bailey (2007). Pests of field crops and pastures: identification and control. CSIRO Publishing. pp. 97–98. ISBN 978-0-643-06758-5.
  37. ^ Roy Van Driesche, Mark Hoddle & Ted Center (2008). Control of pests and weeds by natural enemies: an introduction to biological control. John Wiley and Sons. ISBN 978-1-4051-4571-8.

Read other articles:

Putri ChristinaPutri Christina tahun 1976Kelahiran(1947-02-18)18 Februari 1947Istana Soestdijk, Baarn, BelandaKematian16 Agustus 2019(2019-08-16) (umur 72)Istana Noordeinde, Den Haag, BelandaWangsaOrange-NassauNama lengkapMaria ChristinaAyahPangeran Bernhard dari Lippe-BiesterfeldIbuJuliana dari BelandaPasanganJorge Pérez y Guillermo ​ ​(m. 1975; c. 1996)​AnakBernardo GuillermoNicolás GuillermoJuliana GuillermoAgamaKatolik Romasblm. Calvi...

 

Investigation DiscoveryDiluncurkan27 Januari 2008PemilikWarner Bros. DiscoverySaluran seindukTLC, Animal Planet, Discovery Channel, Travel Channel, Discovery Times, Discovery Kids, The Science Channel, Discovery Home, Military Channel, Discovery Wings, Discovery Civilisation, Discovery Real Time, Discovery Home & Health, Discovery en Español, Discovery Travel & Living, Discovery HD TheaterSitus webinvestigation.discovery.com Investigation Discovery adalah saluran milik Warner Bros. D...

 

American politician (1888–1936) This article needs additional citations for verification. Please help improve this article by adding citations to reliable sources. Unsourced material may be challenged and removed.Find sources: Horace Stansel – news · newspapers · books · scholar · JSTOR (October 2014) (Learn how and when to remove this template message) Horace Stanselc. 192453rd Speaker of the Mississippi House of RepresentativesIn officeJanuary 7, 1...

Artikel Baru Izinkan saya menulis artikel biografi Prof. Dr. H. Abu Yasid, M.A., LL.M karena ia memiliki pranala dan link cukup banyak dan dapat dipertanggung jawabkan secara ilmiah dan secara akademik. Tati Sahamu (bicara) 15 Maret 2022 11.13 (UTC)[balas] Aktive? @Palladin911: Gimana nih? Aktivitas ProyekWiki ini masih kosong. Bahkan untuk anggotanya saja tidak menentu dan tidak jelas. Masalahnya, banyak artikel biografi yang memerlukan penilaian NA. ▪ ꧋ꦩꦣꦪ Fazoffic ( ʖ╎ᓵᔑ�...

 

German general (1899–1970) This article needs additional citations for verification. Please help improve this article by adding citations to reliable sources. Unsourced material may be challenged and removed.Find sources: Fritz Bayerlein – news · newspapers · books · scholar · JSTOR (October 2012) (Learn how and when to remove this template message) Fritz BayerleinBorn(1899-01-14)14 January 1899Würzburg, German EmpireDied30 January 1970(1970-01-30)...

 

In this Arabic name, the surname is Al Nahyan. Sheikh Hazza bin Sultan Al NahyanSheikh ٱلشَّيْخَBorn1905 (according to British sources)[1]DiedJanuary 20, 1958Spouse Maryam bint Said bin Zayed bin Khalifa Al Nahyan Maryam bint Hamdan bin Zayed bin Khalifa Al Nahyan HouseAl NahyanFatherSultan bin Zayed bin Khalifa Al NahyanMotherSalama bint Butti Al-QubaisiReligionIslam Sheikh Hazza bin Sultan Al Nahyan (Arabic: ٱلشَّيْخ هَزَّاع بِن سُلْطَان آل نَ�...

Virginio Gayda Virginio Gayda (Roma, 12 agosto 1885 – Roma, 14 marzo 1944) è stato un giornalista e saggista italiano, direttore dei quotidiani Il Messaggero e Il Giornale d'Italia. Indice 1 Biografia 2 Opere 3 Note 4 Voci correlate 5 Altri progetti 6 Collegamenti esterni Biografia Un numero del Giornale d'Italia diretto da Gayda dal 1926 al 1943 Laureatosi all'Università di Torino in scienze economiche, fu giornalista de La Stampa di Torino dal 1908 e corrispondente da Costantinopoli e v...

 

Zytglogge menjadi markah tanah kota Bern Zytglogge adalah menara jam yang terletak di Bern atau Berne, ibu kota Swiss.[1] Menara ini menjadi salah satu ciri khas kota yang dijuluki sebagai Kota Tua Bern ini dan termasuk Situs Warisan Dunia UNESCO.[2] Sejarah Sebelumnya, menara jam ini merupakan menara pengawas yang beralih fungsi menjadi penjara hingga menjadi menara pengawas kebakaran. Pada tahun 1405, terjadi kebakaran hebat yang mengubah fungsi menara tersebut. Pasca kebaka...

 

JamaikaJamaica (Inggris) Bendera Lambang Semboyan: Out of Many, One People(Inggris: Dari Sekian Banyak, Hanya Satu Rakyat)Lagu kebangsaan:  Jamaica, Land We Love (Indonesia: Jamaika, Tanah Kami tercinta) Lagu kerajaan:  God Save the King (Indonesia: Tuhan Menjaga sang Raja) Perlihatkan BumiPerlihatkan peta BenderaIbu kota(dan kota terbesar)Kingston17°17′N 76°35′W / 17.283°N 76.583°W / 17.283; -76.583Bahasa resmiInggrisBahasa nasionalPatwa Jamaika...

Uranil seng asetat Nama Nama IUPAC seng;dioksouranium(2+);tetraasetat Nama lain seng uranil asetat Penanda Nomor CAS 33959-50-1 N Model 3D (JSmol) Gambar interaktif 3DMet {{{3DMet}}} ChemSpider 59696127 Nomor EC PubChem CID 22673297 Nomor RTECS {{{value}}} InChI InChI=1S/4C2H4O2.2O.U.Zn/c4*1-2(3)4;;;;/h4*1H3,(H,3,4);;;;/q;;;;;;;+4/p-4Key: GWVOTLSLVURWPV-UHFFFAOYSA-J SMILES O=C(C)[O-].[O+]#[U]#[O+].O=C(C)[O-].O=C(C)[O-].[Zn+2].O=C(C)[O-] Sifat Rumus kimia ZnUO2(CH3COO)4 Massa ...

 

1962 compilation album by Perry ComoMr. PresidentCompilation album by Perry ComoReleasedDecember 1962RecordedSeptember 30, October 1, 2, 1962GenreVocalLabelRCA VictorProducerHugo & LuigiPerry Como chronology By Request(1962) Mr. President(1962) The Songs I Love(1963) The Best of Irving Berlin's Songs from Mr. President is a 1962 album by Perry Como, his tenth RCA Victor 12 long-play album.[1] In this album, Como performs select songs from Irving Berlin's then new (and fina...

 

Collegiate basketball team UT Arlington Mavericks 2023–24 UT Arlington Mavericks men's basketball team UniversityUniversity of Texas at ArlingtonHead coachK. T. Turner (1st season)ConferenceWACLocationArlington, TexasArenaCollege Park Center (Capacity: 7,000)NicknameMavericksColorsRoyal blue, white, and orange[1]     Uniforms Home Away NCAA tournament appearances2008Conference tournament champions2008Conference regular season champions2004, 20...

هذه المقالة تحتاج للمزيد من الوصلات للمقالات الأخرى للمساعدة في ترابط مقالات الموسوعة. فضلًا ساعد في تحسين هذه المقالة بإضافة وصلات إلى المقالات المتعلقة بها الموجودة في النص الحالي. (نوفمبر 2023) هذه المقالة يتيمة إذ تصل إليها مقالات أخرى قليلة جدًا. فضلًا، ساعد بإضافة وصلة...

 

Обложка «Деяний» Собора Льво́вский собо́р, также Львовский псевдособор— Собор клириков и мирян Украинской грекокатолической (униатской) церкви (УГКЦ), созванный 8—10 марта 1946 года во Львове, на котором были провозглашены ликвидация Брестской унии 1596 года и присоединени�...

 

2016年美國總統選舉 ← 2012 2016年11月8日 2020 → 538個選舉人團席位獲勝需270票民意調查投票率55.7%[1][2] ▲ 0.8 %   获提名人 唐納·川普 希拉莉·克林頓 政党 共和黨 民主党 家鄉州 紐約州 紐約州 竞选搭档 迈克·彭斯 蒂姆·凱恩 选举人票 304[3][4][註 1] 227[5] 胜出州/省 30 + 緬-2 20 + DC 民選得票 62,984,828[6] 65,853,514[6]...

此條目可能包含不适用或被曲解的引用资料,部分内容的准确性无法被证實。 (2023年1月5日)请协助校核其中的错误以改善这篇条目。详情请参见条目的讨论页。 各国相关 主題列表 索引 国内生产总值 石油储量 国防预算 武装部队(军事) 官方语言 人口統計 人口密度 生育率 出生率 死亡率 自杀率 谋杀率 失业率 储蓄率 识字率 出口额 进口额 煤产量 发电量 监禁率 死刑 国债 ...

 

Ekonomi terbesar di dunia berdasarkan PDB (KKB) pada 2022menurut estimasi Dana Moneter Internasional (dalam triliun dolar internasional) PDB (KKB) berdasarkan negara pada 2022 menurut DMI Berikut adalah daftar negara-negara di dunia berdasarkan produk domestik bruto (PDB) KKB, nilai seluruh produk dan jasa yang dihasilkan di sebuah negara dalam suatu tahun. Perkiraan PDB di sini (dalam dolar internasional) diperoleh dari perhitungan keseimbangan kemampuan berbelanja (KKB). Daftar Daftar menu...

 

本條目存在以下問題,請協助改善本條目或在討論頁針對議題發表看法。 此條目需要編修,以確保文法、用詞、语气、格式、標點等使用恰当。 (2013年8月6日)請按照校對指引,幫助编辑這個條目。(幫助、討論) 此條目剧情、虛構用語或人物介紹过长过细,需清理无关故事主轴的细节、用語和角色介紹。 (2020年10月6日)劇情、用語和人物介紹都只是用於了解故事主軸,輔助�...

American politician For the mayor of St. Louis, Missouri, see John D. Daggett. This article needs additional citations for verification. Please help improve this article by adding citations to reliable sources. Unsourced material may be challenged and removed.Find sources: John Daggett – news · newspapers · books · scholar · JSTOR (September 2014) (Learn how and when to remove this message) John DaggettThe San Francisco Call, June 15, 189616th Lieutena...

 

1963 conflict between Algeria and Morocco Not to be confused with war sand or Sand Wars. Sand WarPart of the Arab Cold War and the Cold WarDateSeptember 25, 1963[4] – February 20, 1964[5] (4 months, 3 weeks and 5 days)LocationAround the oasis towns of Tindouf and FiguigResult Military stalemate[6] The closing of the border south of Figuig, Morocco/Béni Ounif, Algeria. Morocco abandoned its attempts to control Béchar and Tindouf after OAU mediati...