Cowiedesmus

Cowiedesmus
Temporal range: Wenlock/Lochkovian
Scientific classification
Kingdom:
Phylum:
Subphylum:
Class:
Subclass:
Superorder:
Order:
Cowiedesmida
Family:
Cowiedesmidae
Genus:
Cowiedesmus

Wilson & Anderson, 2004
Type species
Cowiedesmus eroticopodus

Cowiedesmus is an extinct millipede genus described from Scotland,[1] and is considered as earliest known land animals alongside Pneumodesmus from same formation.[2] It is originally considered that it is from the middle Silurian, a 2017 study suggested that the geological formation it contains actually appears to be from the Early Devonian (Lochkovian) instead.[3] Although a 2023 study confirmed the age identification of the 2004 study through palynological, palaeobotanical and ziron analyses incorporating newly discovered additional data,[4] this is based on adjacent structurally separated block with different stratigraphy and sedimentology to the block with fossil site it was discovered, and it is confirmed as unsustainable.[5] Cowiedesmus was about 4 cm (1.6 in) long and characterized by a greatly enlarged pair of legs on the 8th segment which may have been used in clasping females or functioned as gonopods (modified legs used to insert sperm during mating). Coweiedesmus is distinct enough from other living and fossil millipedes to be placed in its own order, Cowiedesmida. The only known species, C. eroticopodus, was described in 2004.[1]

The name Cowiedesmus references the Cowie Formation, a geological formation at Cowie Harbor, near the village of Stonehaven, Scotland, where the fossil was found and desmus, a common root word in millipede nomenclature meaning "bond" or "bridge". The specific epithet eroticopodus means "erotic foot".[1]

References

  1. ^ a b c Wilson, Heather M.; Anderson, Lyall I. (2004). "Morphology and taxonomy of Paleozoic millipedes (Diplopoda: Chilognatha: Archipolypoda) from Scotland". Journal of Paleontology. 78 (1): 169–184. doi:10.1666/0022-3360(2004)078<0169:MATOPM>2.0.CO;2.
  2. ^ Russell Garwood & Gregory Edgecombe (2011). "Early terrestrial animals, evolution and uncertainty". Evolution: Education and Outreach. 4 (3): 489–501. doi:10.1007/s12052-011-0357-y.
  3. ^ Suarez, Stephanie E.; Brookfield, Michael E.; Catlos, Elizabeth J.; Stöckli, Daniel F. (28 June 2017). "A U-Pb zircon age constraint on the oldest-recorded air-breathing land animal". PLOS ONE. 12 (6): e0179262. doi:10.1371/journal.pone.0179262. ISSN 1932-6203. PMC 5489152. PMID 28658320.
  4. ^ Wellman, C.H.; Lopes, G.; McKellar, Z.; Hartley, A. (2023). "Age of the basal 'Lower Old Red Sandstone' Stonehaven Group of Scotland: The oldest reported air-breathing land animal is Silurian (late Wenlock) in age". Journal of the Geological Society. The Geological Society of London. doi:10.1144/jgs2023-138. hdl:2164/22754. ISSN 0016-7649.
  5. ^ Brookfield, M. E.; Catlos, E. J.; Garza, H. (7 July 2024). "The oldest 'millipede'-plant association? Age, paleoenvironments and sources of the Silurian lake sediments at Kerrera, Argyll and Bute, Scotland". Historical Biology: 1–13. doi:10.1080/08912963.2024.2367554. ISSN 0891-2963.


Read other articles:

Road in Malaysia Federal Route 3Wakaf Bharu-Kota Bharu-Kubang Kerian Highway(WKK Highway)Route informationPart of AH18 Existed2008–presentHistoryCompleted in 2012Major junctionsNorthwest endWakaf Bharu (East)Pasir PekanMajor intersections FT 134 Jalan Pengkalan Kubur FT 8 Federal Route 8 FT 57 Jalan Long Yunus FT 209 Jalan Pasir Hor FT 131 Jalan Raja Perempuan Zainab II FT 211 Jalan Kubang Kerian-BachokSoutheast endKubang Kerian LocationCountryMalaysiaPrimarydestinationsKota Bharu...

 

Canadian-Palestinian medical doctor Izzeldin AbuelaishBornFebruary 3, 1955Jabalia Camp, Gaza StripNationalityPalestinianEducationUniversity of Cairo MDUniversity of London OB/GynHarvard School of Public Health MPHOccupation(s)Doctor, professor, author, non-profit founderNotable workI Shall Not Hate: A Gaza Doctor's Journey on the Road to Peace and Human DignityTitleMichael and Amira Dan Professor in Global HealthBoard member ofDaughters for Life FoundationWebsitedaughtersforlife.com Izze...

 

Vidkun QuislingPotret Quisling, sekitar tahun 1919 Presiden Menteri NorwegiaMasa jabatan1 Februari 1942 – 9 Mei 1945Menjabat bersama Reichskommissar Josef Terboven PendahuluJohan Nygaardsvold(sebagai Perdana Menteri)PenggantiJohan Nygaardsvold(sebagai Perdana Menteri)Menteri PertahananMasa jabatan12 Mei 1931 – 3 Maret 1933Perdana MenteriPeder KolstadJens Hundseid PendahuluTorgeir Anderssen-RysstPenggantiJens Isak KobroFører of the National UnionMasa jabatan13 Me...

Pour les articles homonymes, voir Lieutenant. Le grade de lieutenant-colonel est un grade d'officier supérieur dans de nombreux pays. Belgique Article connexe : Grades de l'Armée belge. En Belgique, le grade de lieutenant-colonel (Luitenant-kolonel en néerlandais) est le second grade des officiers supérieurs, au-dessus du grade de Major et en dessous de celui de colonel. L'insigne du lieutenant-colonel est constitué d'une barrette et de deux molettes d'éperons héraldique de teint...

 

Autrans-Méaudre en Vercors Autrans. Administration Pays France Région Auvergne-Rhône-Alpes Département Isère Arrondissement Grenoble Intercommunalité Communauté de communes du massif du Vercors Maire Mandat Hubert Arnaud 2020-2026 Code postal 38112, 38880 Code commune 38225 Démographie Populationmunicipale 3 075 hab. (2021) Densité 39 hab./km2 Géographie Coordonnées 45° 07′ 40″ nord, 5° 31′ 40″ est Altitude Min. 940 mMax....

 

Engineering process for electronic hardware This article includes a list of general references, but it lacks sufficient corresponding inline citations. Please help to improve this article by introducing more precise citations. (March 2019) (Learn how and when to remove this template message) Layout view of a simple CMOS Operational Amplifier (inputs are to the left and the compensation capacitor is to the right). The metal layer is coloured blue, green and brown are N- and P-doped Si, the pol...

Philippe Villiers de L'Isle-Adam Mahaguru Ordo Santo YohanesMasa jabatan1521 – 21 Agustus 1534Penguasa monarkiRaja Charles IIPendahuluFabrizio del CarrettoPenggantiPiero de Ponte Informasi pribadiLahir1464Beauvais, PrancisMeninggal21 Agustus 1534 (usia 69–70)Rabat, Hospitaller MaltaMakamKapel St Anne, Benteng St Angelo, Birgu, kemudian dikebumikan ulang di Konkatedral Santo Yohanes, VallettaKebangsaanPrancisKarier militerPihak Ordo Santo YohanesPertempuran/perangPengepungan R...

 

American college basketball season 2016–17 UTSA Roadrunners men's basketballConferenceConference USARecord14–19 (8–10 C-USA)Head coachSteve Henson (1st season)Assistant coaches Mike Peck Scott Thompson Adam Hood Home arenaConvocation CenterSeasons← 2015–162017–18 → 2016–17 Conference USA men's basketball standings vte Conf Overall Team W   L   PCT W   L   PCT Middle Tennessee † 17 – 1   .944 31 – 5   .861 ...

 

Form of pocket billiards popular in Eastern Europe Not to be confused with the historical pocket billiards game pyramid pool, a similar but unrelated game, or slosh or bar billiards, both also called Russian billiards. Russian pyramid game setup, with the object balls in a triangle rack at the foot of the table, and the cue ball in front of (up-table of) the head string (baulk line). Russian pyramid, also known as Russian billiards (Russian: ру́сский билья́рд, russky bilyard),...

Kirtimuka di atas pintu masuk suatu kuil di Kathmandu, Nepal. Kirtimukha (Dewanagari: कीर्तिमुख; ,IAST: kīrtimukha,; arti: muka yang megah) adalah nama makhluk mitologis berwajah monster, dengan taring panjang, mulut menganga, dan lazim ditampilkan sebagai ukiran dalam arsitektur kuil Hindu di Nepal, India, dan Asia Tenggara, dan juga di sejumlah arsitektur Buddhis.[1] Tidak seperti makhluk mitologi Hindu lainnya, seperti monster laut Makara, kirtimuk...

 

2020年夏季奥林匹克运动会波兰代表團波兰国旗IOC編碼POLNOC波蘭奧林匹克委員會網站olimpijski.pl(英文)(波兰文)2020年夏季奥林匹克运动会(東京)2021年7月23日至8月8日(受2019冠状病毒病疫情影响推迟,但仍保留原定名称)運動員206參賽項目24个大项旗手开幕式:帕维尔·科热尼奥夫斯基(游泳)和马娅·沃什乔夫斯卡(自行车)[1]闭幕式:卡罗利娜·纳亚(皮划艇)&#...

 

Доходи населення — грошові кошти і матеріальні блага, які отримує населення для підтримання фізичного, морального, економічного та інтелектуального стану. Найважливішу роль у доходах населення України відіграють оплата праці, на яку у 2014 році припадало 39,9 % загаль...

خنمت نفر حجيت الثانية معلومات شخصية تاريخ الميلاد القرن 19 ق.م  تاريخ الوفاة 2 ألفية ق.م  مكان الدفن دهشور  الزوج سنوسرت الثالث  عائلة الأسرة المصرية الثانية عشر  الحياة العملية المهنة سياسية  تعديل مصدري - تعديل   خنمت نفر حجيت ورت الثانية في الهيروغليفية خ�...

 

يفتقر محتوى هذه المقالة إلى الاستشهاد بمصادر. فضلاً، ساهم في تطوير هذه المقالة من خلال إضافة مصادر موثوق بها. أي معلومات غير موثقة يمكن التشكيك بها وإزالتها. (سبتمبر 2023) تحتاج هذه المقالة إلى تهذيب لتتناسب مع دليل الأسلوب في ويكيبيديا. فضلاً، ساهم في تهذيب هذه المقالة من خلا...

 

Overview of driving in Singapore This article needs additional citations for verification. Please help improve this article by adding citations to reliable sources. Unsourced material may be challenged and removed.Find sources: Driving in Singapore – news · newspapers · books · scholar · JSTOR (July 2020) (Learn how and when to remove this message) The Pan Island Expressway, one of the main expressways in the Singapore road network In Singapore, cars a...

Arena in Sydney, New South Wales, Australia For the old Sydney Showground located in Moore Park, see Sydney Showground (Moore Park). Arriving at the Showground during the Royal Easter Show The Sydney Showground is a purpose-built venue used each year for the Sydney Royal Easter Show. Located at Sydney Olympic Park in Sydney, New South Wales, Australia, it was opened in 1998, as a venue for the 2000 Summer Olympics and to replace the former Sydney Showground at Moore Park. Sydney Showground is...

 

Meryl Streep, Hon. D.A. 2010[1] (l-r) US Navy Secretary (and Har­vard Over­seer) George Meyer; Prof. Frank W. Taussig; Pres. Eliot (who dis­dained aca­dem­ic regal­ia);[2] Bishop (and Over­seer) William Lawrence (1911) What was originally called Harvard Colledge[3] (around which Harvard University eventually grew)[4] held its first Commence­ment in September 1642, when nine degrees were conferred.[5] Today some 1...

 

Jacques Stany nel film L'arcangelo (1969) Jacques Stany, pseudonimo di Jacob Stanislawski (Minsk, 4 aprile 1930[1] – Roma, 19 luglio 2016), è stato un attore e doppiatore bielorusso naturalizzato francese. Ha recitato per la maggior parte della sua carriera in Italia. Indice 1 Biografia 2 Filmografia 2.1 Cinema 2.2 Televisione 3 Doppiaggio 4 Doppiatori italiani 5 Note 6 Altri progetti 7 Collegamenti esterni Biografia Jacques Stany è ricordato in Italia soprattutto per le sue parte...

American art historian Portrait of Else Glaser (wife of Curt Glaser) by Edvard Munch Curt Glaser (May 29, 1879 (Leipzig) – November 23, 1943 (Lake Placid, New York, USA)) was a German Jewish art historian, art critic and collector who was persecuted by the Nazis.[1] Life Portrait of Max Beckmann (1929) Gedenktafel, Matthäikirchplatz 8, in Berlin-Tiergarten Glaser's parents, the businessman Simon Glaser (1841–1904) and his wife Emma Glaser, née Haase (1854–1927), moved to Berli...

 

Theatre in Surrey, England Sketch of the interior of the Richmond Theatre Exterior of the Richmond Theatre and Edmund Kean's House. Kean was manager of the Richmond Theatre from 1831 until his death in 1833. The Richmond Theatre was a theatre located in Richmond, Surrey, England. It was built in 1765 under the supervision of David Garrick. It was one of Surrey's major theatres for nearly 120 years.[1] The theatre was originally named The New Theatre on Richmond Greene and opened on 15...