^ 3.03.1van der Kuyl, Antoinette C.; Ballasina, Donato L. Ph.; Dekker, John T.; Maas, Jolanda; Willemsen, Ronald E. & Goudsmit, Jaap. Phylogenetic Relationships among the Species of the Genus Testudo (Testudines: Testudinidae) Inferred from Mitochondrial 12S rRNA Gene Sequences. Molecular Phylogenetics and Evolution. 2002, 22 (2): 174–183. doi:10.1006/mpev.2001.1052.
^Uwe Fritza, Pavel Široký, Hajigholi Kamic and Michael Wink. Environmentally caused dwarfism or a valid species—Is Testudo weissingeri Bour, 1996 a distinct evolutionary lineage? New evidence from mitochondrial and nuclear genomic markers. Molecular Phylogenetics and Evolution. 2005, 37 (2): 389–401. doi:10.1016/j.ympev.2005.03.007.
^Široký, Pavel & Fritz, Uwe. Is Testudo werneri a distinct species?. Biologia. 2007, 62 (2): 228–231. doi:10.2478/s11756-007-0036-1.
^da Nóbrega Alves, Rômulo Romeu; da Silva Vieira; Washington Luiz & Gomes Santana, Gindomar. Reptiles used in traditional folk medicine: conservation implications. Biodiversity and Conservation. 2008, 17 (8): 2037–2049. doi:10.1007/s10531-007-9305-0.