↑U.S. Johansson, J. Ekman, R.C.K. Bowie, P. Halvarsson, J.I. Ohlson, T.D. Price et P.G.P. Ericson, « A complete multilocus species phylogeny of the tits and chickadees (Aves: Paridae) », Molecular Phylogenetics and Evolution, vol. 69, no 3, , p. 852–860 (PMID23831453, DOI10.1016/j.ympev.2013.06.019)