The squadron formed at RAF Hutton Cranswick in Yorkshire on 20 November 1941. It was equipped initially with Miles Martinets and then Hawker Hurricanes and Lockheed Hudsons. Targets were towed by aircraft to provide practice for the anti-aircraft defences in Eastern England. The squadron was also equipped with the Vultee Vengeance before it was disbanded on 26 June 1945.