2012 in paleoentomology
Overview of the events of 2012 in paleoentomology
List of years in paleoentomology
This list of 2012 in paleoentomology records new fossil insect taxa that are to be described during the year, as well as documents significant paleoentomology discoveries and events which occurred during that year.
Blattodea
Coleoptera
Name
Novelty
Status
Authors
Age
Unit
Location
Notes
Images
Abrocar concavus [ 12]
Sp. nov
Valid[ 13]
Davis et al.
Early Cretaceous
Yixian Formation
China
A weevil , a species of Abrocar .
Abrocar relicinus [ 12]
Sp. nov
Valid[ 13]
Davis et al.
Early Cretaceous
Yixian Formation
China
A weevil , a species of Abrocar .
Abrohadeocoleodes [ 14]
Gen. et 4 sp. nov
Valid[ 15]
Tan et al.
Middle Jurassic
Jiulongshan Formation
China
A schizophorid archostematan beetle . Genus contains four species: A. eurycladus , A. ooideus , A. nii and A. patefactus .
Alloioscarabaeus [ 16]
Gen. et sp. nov
Valid
Bai, Ren & Yang in Bai et al.
Middle Jurassic
Jiulongshan Formation
China
An alloioscarabaeid scarabaeoid beetle . The type species is Alloioscarabaeus cheni .
Alphitopsis [ 17]
Gen. et sp. nov
Valid
Kirejtshuk, Nabozhenko & Nel
Early Cretaceous
Yixian Formation
China
A tenebrionine darkling beetle . The type species is Alphitopsis initialis .
Anotylus archaicus [ 18]
sp nov
Valid
Yue, Makranczy & Ren
Early Cretaceous
Yixian Formation
China
An oxyteline rove beetle , a species of Anotylus .
Aphebodactyla [ 19]
Gen. et sp. nov
Valid
Chatzimanolis et al.
Cretaceous
Burmese amber
Myanmar
A ptilodactylid byrrhoid beetle. The type species is Aphebodactyla rhetine .
Aploceble (Chalcoaploceble) viridiaeneus [ 20]
Subgen. et sp. nov
Valid
Tshernyshev
Eocene
Baltic amber
Russia (Kaliningrad Oblast )
A melyrid beetle belonging to the subfamily Dasytinae , a subgenus and species of Aploceble .
Apticax [ 21]
Gen. et 2 sp. nov
Valid
Schomann & Solodovnikov
Early Cretaceous (Aptian–Albian )
Crato Formation
Brazil
A rove beetle of uncertain phylogenetic placement. The type species is A. volans ; genus also includes A. solidus .
Archaeosciaphilus [ 22]
Gen. et sp. nov
Valid
Legalov
Eocene
Europe
A curculionid beetle found in Baltic amber . The type species is Archaeosciaphilus marshalli .
Attagenus yantarnyi [ 23]
Sp. nov
Valid
Háva & Bukejs
Late Eocene
Prussian Formation
Russia (Kaliningrad Oblast )
A dermestid beetle found in Baltic amber , a species of Attagenus .
Baltocar groehni [ 24]
Sp. nov
Valid
Riedel in Riedel et al.
Eocene
Baltic amber
Russia ( Kaliningrad Oblast )
A weevil belonging to the family Attelabidae .
Baltocar hoffeinsorum [ 24]
Sp. nov
Valid
Riedel in Riedel et al.
Eocene
Baltic amber
Russia ( Kaliningrad Oblast )
A weevil belonging to the family Attelabidae.
Baltocar subnudus [ 24]
Sp. nov
Valid
Riedel in Riedel et al.
Eocene
Baltic amber
Russia ( Kaliningrad Oblast )
A weevil belonging to the family Attelabidae.
Cartodere (Aridius) succinobaltica [ 25]
Sp. nov
Valid
Bukejs, Rücker & Kirejtshuk
Late Eocene
Europe
A latridiid beetle found in Baltic amber , a species of Cartodere .
Catops perkovskyi [ 26]
Sp. nov
Valid
Perreau
Eocene
Russia (Kaliningrad Oblast )
A leiodid beetle found in Baltic amber , a species of Catops .
Conapium (Palaeoconapion) alleni [ 27]
Subgen. et sp. nov
Valid
Legalov
Eocene
Baltic amber
Russia (Kaliningrad Oblast )
A beetle belonging to the family Brentidae , a subgenus and species of Conapium .
Crassisorus [ 28]
Gen. et sp. nov
Valid
Nikolajev, Wang & Zhang
Early Cretaceous
Yixian Formation
China
A member of the family Hybosoridae . The type species is C. fractus .
Cretohypna [ 29]
Gen. et sp. nov
Valid
Yan, Nikolajev & Ren
Early Cretaceous
Yixian Formation
China
A glaphyrid beetle . The type species is Cretohypna cristata .
Cretotanaos [ 30]
Gen. et sp. nov
Valid[ 31]
Legalov
Early Cretaceous (Aptian)
Mongolia
An apionine brentid weevil . Type species is Cretotanaos bontsaganensis .
Cryptophagus harenus [ 32]
Sp. nov
Valid
Lyubarsky & Perkovsky
Late Eocene
Ukraine
A cryptophagid beetle found in Rovno amber , a species of Cryptophagus .
Electotreta [ 33]
Gen. et sp.
Valid
Kazantsev
Eocene
Lithuania
A firefly . The type species is Electotreta rasnitsyni .
Electromethes [ 33]
Gen. et sp. nov
Valid
Kazantsev
Eocene
Lithuania
An omethid beetle. The type species is Electromethes alleni .
Electropteron [ 34]
Gen. et sp. nov
Valid
Kazantsev
Oligocene
Dominican Republic
A lycid . The type species is E. avus .
Elodes beigeli [ 35]
Sp. nov
Valid
Klausnitzer
Eocene
Europe
A scirtid beetle found in Baltic amber , a species of Elodes .
Elodes mysticopalpalis [ 35]
Sp. nov
Valid
Klausnitzer
Eocene
Europe
A scirtid beetle found in Baltic amber , a species of Elodes .
Enicmus adrianae [ 36]
Sp. nov
Valid
Bukejs, Reike & Rücker
Late Eocene
Russia (Kaliningrad Oblast )
A member of Latridiidae found in Baltic amber , a species of Enicmus .
Eocenorhynchites [ 22]
Gen. et sp. nov
Valid
Legalov
Eocene
Europe
A rhynchitid beetle found in Baltic amber . The type species is Eocenorhynchites vossi .
Eodromeus daohugouensis [ 37]
sp nov
Valid
Wang, Zhang and Ponomarenko
Middle Jurassic
Daohugou Beds
China
A trachypachid , a species of Eodromeus .
Eodromeus robustus [ 37]
sp nov
Valid
Wang, Zhang and Ponomarenko
Middle Jurassic
Daohugou Beds
China
A trachypachid, a species of Eodromeus .
Eoluciola [ 38]
Gen. et sp. nov
Valid
Kazantsev
Eocene
Europe (Baltic Sea coast)
A lucioline firefly found in Baltic amber . The type species is Eoluciola varang .
Euroleptochromus [ 39]
Gen. et sp. nov
Valid
Jałoszyński
Eocene
Lithuania
A rove beetle found in Lithuanian Eocene Baltic amber . The type species is Euroleptochromus sabathi .
Helophorus (Mesosperchus) inceptivus [ 40]
Sp. nov
Valid
Fikáček et al.
Late Jurassic
Mongolia
A hydrophiloid beetle , a species of Helophorus .
Helophorus (Mesosperchus) yixianus [ 40]
Sp. nov
Valid
Fikáček et al.
Jurassic-Cretaceous boundary
Yixian Formation
China
A hydrophiloid beetle , a species of Helophorus .
Hydroscapha? jeholensis [ 41]
Sp. nov
Valid
Cai, Short & Huang
Early Cretaceous
Yixian Formation
China
A member of the family Hydroscaphidae .
Juroglypholoma [ 42]
Gen. et sp. nov
Valid
Cai et al.
Middle Jurassic
Daohugou Beds
China
A glypholomatine rove beetle . The type species is Juroglypholoma antiquum .
Kararhynchus gratshevi [ 43]
Species
Valid
Legalov
Middle-Late Jurassic
Karabastau Formation
Kazakhstan
An obrieniid beetle , a species of Kararhynchus .
Kararhynchus jurassicus [ 43]
Species
Valid
Legalov
Middle-Late Jurassic
Karabastau Formation
Kazakhstan
An obrieniid beetle, a species of Kararhynchus .
Laetopsia [ 40]
Gen. et comb. nov
Valid
Fikáček et al.
Early Cretaceous
China
Mongolia
Russia
A hydrophiloid beetle ; a new genus for "Hydrophilopsia " baissensis Ponomarenko (1987), "H." bontsaganica Prokin (2009), "H." hydraenoides Prokin, Ren & Fikáček (2010), "H." mongolica Ponomarenko (1987) and "H." shatrovskiyi Prokin, Ren & Fikáček (2010).
Lasiosyne cataphracta [ 44]
Sp. nov
Valid
Yan
Early Cretaceous
Mongolia
A lasiosynid beetle , a species of Lasiosyne .
Lasiosyne decora [ 44]
Sp. nov
Valid
Yan
Early Cretaceous
Russia
A lasiosynid beetle , a species of Lasiosyne .
Lasiosyne insculpta [ 44]
Sp. nov
Valid
Yan
Late Jurassic
Mongolia
A lasiosynid beetle , a species of Lasiosyne .
Lasiosyne longitarsa [ 44]
Sp. nov
Valid
Yan
Late Jurassic
Mongolia
A lasiosynid beetle , a species of Lasiosyne .
Lasiosyne punctata [ 44]
Sp. nov
Valid
Yan
Early Cretaceous
Mongolia
A lasiosynid beetle , a species of Lasiosyne .
Lasiosyne shartegiensis [ 44]
Sp. nov
Valid
Yan
Late Jurassic
Mongolia
A lasiosynid beetle , a species of Lasiosyne .
Leehermania [ 45]
Gen. et sp. nov
Valid
Chatzimanolis, Grimaldi & Engel in Chatzimanolis et al.
Late Triassic (late Carnian or early Norian)
Cow Branch Formation
United States
A beetle of uncertain phylogenetic placement, might be the earliest known rove beetle or a member of Myxophaga related to the family Hydroscaphidae .[ 46] The type species is Leehermania prorova .
Lithohypna lepticephala [ 47]
Sp. nov
Valid
Nikolajev & Ren
Early Cretaceous
Yixian Formation
China
A beetle belonging to the family Glaphyridae , a species of Lithohypna .
Lithohypna longula [ 47]
Sp. nov
Valid
Nikolajev & Ren
Early Cretaceous
Yixian Formation
China
A beetle belonging to the family Glaphyridae , a species of Lithohypna .
Lithohypna tuberculata [ 47]
Sp. nov
Valid
Nikolajev & Ren
Early Cretaceous
Yixian Formation
China
A beetle belonging to the family Glaphyridae , a species of Lithohypna .
Lithohypna yuxiana [ 47]
Sp. nov
Valid
Nikolajev & Ren
Early Cretaceous
Yixian Formation
China
A beetle belonging to the family Glaphyridae , a species of Lithohypna .
Macratria alleni [ 48]
Sp. nov
Valid
Telnov
Late Eocene
Latvia or Lithuania
A macratriine anthicid beetle found in Baltic amber , a species of Macratria .
Melanapion (Melanapionoides) wanati [ 27]
Subgen. et sp. nov
Valid
Legalov
Eocene
Baltic amber
Russia (Kaliningrad Oblast )
A beetle belonging to the family Brentidae , a subgenus and species of Melanapion .
Menopraesagus oryziformis [ 14]
Sp. nov
Valid[ 49]
Tan et al.
Middle Jurassic
Jiulongshan Formation
China
A schizophorid archostematan beetle , a species of Menopraesagus .
Nephus subcircularis [ 50]
Sp. nov
Valid
Kirejtshuk & Nel
Earliest Eocene
France
A scymnine coccinellid beetle found in Oise amber, as species of Nephus
Notocupes lini [ 51]
Sp. nov
Valid
Ponomarenko et al.
Early Jurassic
China
Originally described as a species of Notocupes ; Kirejtshuk (2020) transferred this species to the genus Zygadenia .[ 52]
Olemehlia [ 53]
Gen. et sp. nov
Junior homonym
Batelka
Eocene (Lutetian )
Baltic amber
Lithuania
A member of the family Ripiphoridae belonging to the subfamily Ripidiinae . The type species is O. krali . The generic name is preoccupied by Olemehlia Holzschuh (2011); Batelka (2017) coined a replacement name Olemehliella .[ 54]
Omma delicata [ 55]
Sp. nov
Tan et al.
Middle Jurassic
Jiulongshan Formation
China
A species of Omma
Orchesia (Orchestera) turkini [ 56]
Sp. nov
Valid
Alekseev & Bukejs
Late Eocene
Prussian Formation
Russia (Kaliningrad Oblast )
A melandryid beetle found in Baltic amber , a species of Orchesia .
Palaeoerirhinus [ 30]
Gen. et 4 sp. nov
Valid[ 31]
Legalov
Early Cretaceous (Aptian)
Mongolia
A curculionid weevil . Type species is Palaeoerirhinus ponomarenkoi ; genus also contains new species P. latus , P. thompsoni and P. longirostris .
Palaeometrioxena [ 22]
Gen. et sp. nov
Junior synonym
Legalov
Eocene
Baltic amber
Europe (Baltic Sea region)
A belid . The type species is Palaeometrioxena zherikhini . Legalov (2019) considered the genus Palaeometrioxena to be a junior synonym of the genus Archimetrioxena , and transferred the species P zherikhini to the latter genus.[ 57]
Pareuryomma [ 55]
Nom. et 2 sp. nov
Tan et al.
Early Cretaceous
Yixian Formation
China
A member of the family Ommatidae ; a replacement name for Euryomma Tan et al. (2006). The type species is "Euryomma" tylodes Tan et al. (2006); genus also includes new species P. cardiobasis . Additional new species described by Tan et al. , P. ancistrodonta from the Middle Jurassic Jiulongshan Formation , was subsequently transferred to the genus Omma by Kirejtshuk (2020).[ 52]
Pleocoma dolichophylla [ 58]
Sp. nov
Valid
Nikolajev & Ren
Early Cretaceous
Yixian Formation
China
A rain beetle , a species of Pleocoma .
Prosinodendron [ 59]
Gen. et sp. nov
Valid
Bai, Ren, & Yang
Barremian to early Aptian
Yixian Formation
China
A stag beetle . The type species is Prosinodendron krelli .
Prosolierius [ 60]
Gen. et 3 sp. nov
Valid
Thayer, Newton, & Chatzimanolis
Barremian to Albian
Lebanon
Myanmar
A rove beetle .
Pseudobrienia [ 43]
Gen. et sp. nov
Valid
Legalov
Middle-Late Jurassic
Karabastau Formation
Kazakhstan
An obrieniid beetle. Its type species is Pseudobrienia rasnitsyni .
Pseudoglaesotropis [ 22]
Gen. et sp. nov
Valid
Legalov
Eocene
Baltic amber
Europe (Baltic Sea region)
An anthribid beetle. The type species is Pseudoglaesotropis martynovi . Legalov (2019) subsequently changed the rank of the genus Pseudoglaesotropis to the subgenus of the genus Glaesotropis , and transferred the species P. martynovi to the latter genus.[ 57]
Pulcherhybosorus [ 61]
Gen. et sp. nov
Valid
Yan, Bai & Ren
Early Cretaceous
Yixian Formation
China
A hybosorid beetle. The type species is Pulcherhybosorus tridentatus .
Rhyzobius antiquus [ 50]
Sp. nov
Valid
Kirejtshuk & Nel
Earliest Eocene
Oise amber
France
A cocciduline coccinellid beetle , a species of Rhyzobius
Rhyzobius gratiosus [ 50]
Sp. nov
Valid
Kirejtshuk & Nel
Earliest Eocene
Oise amber
France
A cocciduline coccinellid beetle, a species of Rhyzobius
Septiventer [ 62]
Gen. et sp. nov
Valid[ 63]
Bai et al.
Late Jurassic or Early Cretaceous
Yixian Formation
China
A septiventerid scarabaeoid beetle . The type species is Septiventer quadridentatus .
Shartegopsis [ 64]
Gen. et sp. nov
Valid
Kirejtshuk
Late Jurassic
Mongolia
A clerid beetle . The type species is Shartegopsis miranda .
Sinodromeus [ 37]
Gen. et sp. nov
Valid
Wang, Zhang & Ponomarenko
Early Cretaceous
Yixian Formation
China
A trachypachid. The type species is Sinodromeus liutiaogouensis .
Sinopeltis [ 65]
Gen. et 2 sp. nov
Valid
Yu et al.
Middle Jurassic
Jiulongshan Formation
China
A peltine trogossitid beetle . Genus contains two species: S. jurrasica (type species) and S. amoena . Kirejtshuk (2017) considered the genus Sinopeltis to be a junior synonym of the genus Cervicatinius Tan & Ren (2007).[ 66]
Sinoschizala [ 67]
Gen. et sp. nov
Valid
Jarzembowski et al.
Middle Jurassic
Daohugou Beds
China
A schizophorid beetle . The type species is Sinoschizala darani .
Succinometrioxena [ 68]
Gen. et sp. nov
Valid
Legalov
Eocene
Europe
A belid weevil found in Baltic amber . The type species is Succinometrioxena poinari .
Sucinoptinus bukejsi [ 69]
Sp. nov
Valid
Alekseev
Eocene (Priabonian )
Prussian Formation
Russia (Kaliningrad Oblast )
A spider beetle found in Baltic amber , a species of Sucinoptinus .
Tafforeus [ 26]
Gen. et sp. nov
Valid
Perreau
Eocene
Russia (Kaliningrad Oblast )
A leiodid beetle found in Baltic amber . The type species is Tafforeus cainosternus .
Talbragarus [ 70]
Gen. et sp. nov
Valid
Oberprieler & Oberprieler
Late Jurassic
Talbragar Fish Bed
Australia
A nemonychid weevil . The type species is Talbragarus averyi .
Tetraphalerus decorosus [ 55]
Sp. nov
Tan et al.
Middle Jurassic
Jiulongshan Formation
China
Originally described as a species of Tetraphalerus ; Kirejtshuk (2020) transferred this species to the ommatine genus Monticupes .[ 52]
Toksunius [ 51]
Gen. et comb. nov
Valid
Ponomarenko et al.
Late Triassic
China
A new genus for "Ademosynoides " chinaticus Lin (1992).
Tomoderus balticus [ 71]
Sp. nov
Valid
Telnov
Late Eocene
Latvia or Lithuania
A tomoderine anthicid beetle found in Baltic amber , a species of Tomoderus .
Trixagus majusculus [ 72]
sp nov
Valid
Kovalev, Kirejtshuk & Nel
Lowermost Eocene
France
A throscid beetle , a species of Trixagus .
Unda chifengensis [ 37]
sp nov
Valid
Wang, Zhang & Ponomarenko
Middle Jurassic
Daohugou Beds
China
A trachypachid, a species of Unda .
Xenohimatium [ 73]
Gen. et sp. nov
Valid
Lyubarsky & Perkovsky
Late Eocene
Rovno amber
Ukraine
A pleasing fungus beetle belonging to the subfamily Xenoscelinae . The type species is X. rovnense .
Diptera
Name
Novelty
Status
Authors
Age
Unit
Location
Notes
Images
Aetheapnomyia [ 74]
Gen. et comb. nov
Valid
Harbach & Greenwalt
Eocene
Europe
A mosquito found in Baltic Amber , a new genus for "Aedes " hoffeinsorum Szadziewski 1998.
Bibio anatolicus [ 75]
Sp. nov
Valid
Skartveit & Nel
Oligocene
Turkey
A species of Bibio .
Bibio nigricosta [ 75]
Sp. nov
Valid
Skartveit & Nel
Oligocene
Turkey
A species of Bibio .
Calosargus talbragarensis [ 76]
Sp. nov
Valid
Oberprieler & Yeates
Late Jurassic
Australia
An archisargid brachyceran , a species of Calosargus .
Clarumreddera [ 77]
Gen. et 2 sp. nov
Valid
Fedotova & Perkovsky
Late Eocene
Rovno amber
Ukraine
A cecidomyiid fly , with two species: C. korneyevi and C. conceptiva .
Culiseta kishenehn [ 74]
Sp. nov
Valid
Harbach & Greenwalt
Eocene
Kishenehn Formation
United States
A mosquito , a species of Culiseta .
Culiseta lemniscata [ 74]
Sp. nov
Valid
Harbach & Greenwalt
Eocene
Kishenehn Formation
United States
A mosquito , a species of Culiseta .
Curtonotum electrodominicum [ 78]
Sp. nov
Valid
Grimaldi & Kirk-Spriggs
Miocene
Dominican Republic
A curtonotid found in Dominican amber , a species of Curtonotum .
Daohugosargus [ 79]
Gen. et comb. nov
Valid
Zhang
Middle Jurassic
Daohugou Beds
China
An archisargid brachyceran ; a new genus for "Sharasargus " eximius Zhang, Yang & Ren (2008).
Depressonotum [ 78]
Gen. et sp. nov
Valid
Grimaldi & Kirk-Spriggs
Miocene
Dominican Republic
A curtonotid found in Dominican amber . The type species is Depressonotum priscum .
Dicranomyia (Caenolimonia) alexbrowni [ 80]
Sp. nov
Valid
Podenas & Poinar
Late Oligocene to early Miocene
Mexico
A species of Dicranomyia .
Dicranomyia (Dicranomyia) chiapa [ 80]
Sp. nov
Valid
Podenas & Poinar
Late Oligocene to early Miocene
Mexico
A species of Dicranomyia .
Dicranomyia (Dicranomyia) mexa [ 80]
Sp. nov
Valid
Podenas & Poinar
Late Oligocene to early Miocene
Mexico
A species of Dicranomyia .
Dicranomyia (Dicranomyia) vella [ 80]
Sp. nov
Valid
Podenas & Poinar
Late Oligocene to early Miocene
Mexico
A species of Dicranomyia .
Didactylomyia dlusskyi [ 81]
Sp. nov
Valid
Fedotova & Perkovsky
Late Eocene
Ukraine
A cecidomyiid fly found in Rovno amber, a species of Didactylomyia .
Enischnomyia [ 82]
Gen. et sp. nov
Valid
Poinar & Brown
Burdigalian
Dominican amber
Dominican Republic
A bat fly, type species E. stegosoma , host for the Plasmodiidae Vetufebrus ovatus [ 83]
Enischnomyia stegosoma
Eoaedes [ 74]
Gen. et comb. nov
Valid
Harbach & Greenwalt
Eocene
Europe
A mosquito found in Baltic Amber , a new genus for "Aedes " damzeni Szadziewski 1998.
Eoptychopterina postica [ 84]
Sp. nov
Valid
Liu, Shih & Ren
Middle Jurassic
Jiulongshan Formation
China
A ptychopterid , a species of Eoptychopterina .
Eotrichocera (Archaeotrichocera) spatiosa [ 84]
Sp. nov
Valid
Liu, Shih & Ren
Middle Jurassic
Jiulongshan Formation
China
A trichocerid , a species of Eotrichocera .
Erioptera (?Mesocyphona) beata [ 80]
Sp. nov
Valid
Podenas & Poinar
Late Oligocene to early Miocene
Mexico
A species of Erioptera .
Erioptera (Mesocyphona) diver [ 80]
Sp. nov
Valid
Podenas & Poinar
Late Oligocene to early Miocene
Mexico
A species of Erioptera .
Erioptera (?Mesocyphona) sherry [ 80]
Sp. nov
Valid
Podenas & Poinar
Late Oligocene to early Miocene
Mexico
A species of Erioptera .
Flagellisargus [ 85]
Gen. et 3 sp. nov
Valid
Zhang
Callovian or Oxfordian
Daohugou Beds
China
An archisargid fly. Genus contains three species: Flagellisargus sinicus , F. venustus and F. robustus .
Groveromyia [ 86]
Gen. et 3 sp. et comb. nov
Valid
Fedotova & Perkovsky
Late Eocene
Rovno amber
Ukraine
A cecidomyiid fly with 4 species G. occlusa ; G. digna , G. astrosa , and "Bryocrypta " concinna Fedotova (2005).
Helius (Helius) collemus [ 80]
Sp. nov
Valid
Podenas & Poinar
Late Oligocene to early Miocene
Mexico
A species of Helius .
Jurochlus adustus [ 87]
Sp. nov
Valid
Lukashevich & Przhiboro
Mesozoic
Mongolia
A chironomid , a species of Jurochlus .
Jurochlus limbatus [ 87]
Sp. nov
Valid
Lukashevich & Przhiboro
Mesozoic
Mongolia
A chironomid , a species of Jurochlus .
Jurochlus lineatus [ 87]
Sp. nov
Valid
Lukashevich & Przhiboro
Mesozoic
Mongolia
A chironomid , a species of Jurochlus .
Jurochlus trivittatus [ 87]
Sp. nov
Valid
Lukashevich & Przhiboro
Mesozoic
Mongolia
A chironomid , a species of Jurochlus .
Lacrimyza [ 88]
Gen. et 2 sp. nov
Valid[ 89]
Roháček
Eocene
Europe
An anthomyzid . Genus contains two species: L. lacrimosa and L. christelae .
Laiyangitabanus [ 90]
Gen. et sp. nov
Valid
Zhang
Early Cretaceous
Laiyang Formation
China
A horse-fly . The type species is Laiyangitabanus formosus .
Lithorhagio [ 91]
Gen. et sp. nov
Valid
Zhang & Li
Middle Jurassic or lowermost Upper Jurassic
Daohugou Beds
China
A rhagionid . The type species is Lithorhagio megalocephalus .
Lygistorrhina caribbiana [ 92]
Sp. nov
Valid
Grund
Miocene
Dominican Republic
A lygistorrhinid fungus gnat found in Dominican amber , a species of Lygistorrhina .
Mesorhyphus handlirschi [ 93]
Sp. nov
Valid
Lukashevich
Upper Jurassic
Mongolia
An anisopodid bibionomorph , a species of Mesorhyphus .
Mesorhyphus hennigi [ 93]
Sp. nov
Valid
Lukashevich
Upper Jurassic
Mongolia
An anisopodid bibionomorph , a species of Mesorhyphus .
Mesosciophila sigmoidea [ 94]
sp nov
Valid
Wang, Zhao & Ren
Middle Jurassic
Jiulongshan Formation
China
A mesosciophilid nematoceran , a species of Mesosciophila .
Mesosolva zhangae [ 79]
Nom. nov
Valid
Zhang
Middle Jurassic
Daohugou Beds
China
An archisargid brachyceran ; a replacement name for Brevisolva daohugouensis Zhang et al. (2010).
Nemopalpus inexpectatus [ 95]
Sp. nov
Valid
Wagner in Wagner & Stuckenberg
Late Eocene
Baltic amber
Europe
A bruchomyiine psychodid fly . Originally described as a species of Nemopalpus , but subsequently transferred to the genus Palaeosycorax .[ 96]
Nemopalpus velteni [ 95]
Sp. nov
Valid
Wagner in Wagner & Stuckenberg
Cretaceous (Albian)
Burmese amber
Myanmar
A bruchomyiine psychodid fly . Originally described as a species of Nemopalpus , but subsequently transferred to the genus Palaeoglaesum .[ 96]
Orientisargus [ 79]
Gen. et sp. nov
Valid
Zhang
Latest Middle Jurassic or earliest Late Jurassic
Daohugou Beds
China
An orientisargid archisargoid brachyceran . The type species is Orientisargus illecebrosus .
Oryctochlus brundini [ 97]
Sp. nov
Valid
Lukashevich
Late Jurassic
Mongolia
A podonomine chironomid , as species of Oryctochlus
Oryctochlus kaluginae [ 97]
Sp. nov
Valid
Lukashevich
Late Jurassic
Mongolia
A podonomine chironomid , as species of Oryctochlus
Palaeopoecilostola eocenica [ 98]
Sp. nov
Valid
Kania & Krzemiński
Eocene
Europe
A limoniid found in Baltic amber , a species of Palaeopoecilostola .
Paramesosciophilodes aequus [ 94]
sp nov
Valid
Wang, Zhao & Ren
Middle Jurassic
Jiulongshan Formation
China
A mesosciophilid nematoceran , a species of Paramesosciophilodes .
Penthetria beskonakensis [ 75]
Sp. nov
Valid
Skartveit & Nel
Oligocene
Turkey
A species of Penthetria .
Probittacomorpha brisaci [ 99]
sp nov
Valid
Krzemiński, Kania & Nel
Late Miocene
France
A ptychopterid , a species of Probittacomorpha .
Procramptonomyia kovalevi [ 93]
Sp. nov
Valid
Lukashevich
Upper Jurassic
Kazakhstan
Mongolia
A procramptonomyiid bibionomorph , a species of Procramptonomyia .
Procramptonomyia ponomarenkoi [ 93]
Sp. nov
Valid
Lukashevich
Upper Jurassic
Mongolia
A procramptonomyiid bibionomorph , a species of Procramptonomyia .
Protanthomyza hennigi [ 88]
Sp. nov
Valid[ 89]
Roháček
Eocene
Europe
An anthomyzid , a species of Protanthomyza .
Protanthomyza hoffeinsorum [ 88]
Sp. nov
Valid[ 89]
Roháček
Eocene
Europe
An anthomyzid , a species of Protanthomyza .
Protanthomyza krylovi [ 88]
Sp. nov
Valid[ 89]
Roháček
Eocene
Europe
An anthomyzid , a species of Protanthomyza .
Protanthomyza loewi [ 88]
Sp. nov
Valid[ 89]
Roháček
Eocene
Europe
An anthomyzid , a species of Protanthomyza .
Protanthomyza meunieri [ 88]
Sp. nov
Valid[ 89]
Roháček
Eocene
Europe
An anthomyzid , a species of Protanthomyza .
Protanthomyza presli [ 88]
Sp. nov
Valid[ 89]
Roháček
Eocene
Europe
An anthomyzid , a species of Protanthomyza .
Protanthomyza tschirnhausi [ 88]
Sp. nov
Valid[ 89]
Roháček
Eocene
Europe
An anthomyzid , a species of Protanthomyza .
Protoculicoides ciliatus [ 100]
Sp. nov
Valid
Borkent
Cretaceous
Canadian amber
Canada
A member of the family Ceratopogonidae . Originally described as a species of Protoculicoides ; Borkent (2019) transferred this species to the genus Atriculicoides .[ 101]
Protorhagio parvus [ 91]
sp nov
Valid
Zhang & Li
Middle Jurassic or lowermost Upper Jurassic
Daohugou Beds
China
A rhagionid, a species of Protorhagio .
Protorhyphus rohdendorfi [ 93]
Sp. nov
Valid
Lukashevich
Upper Jurassic
Mongolia
A protorhyphid bibionomorph , a species of Protorhyphus .
Rovnodidactylomyia [ 81]
Gen. et 3 sp. et comb. nov
Valid
Fedotova & Perkovsky
Late Eocene
Europe
A cecidomyiid fly with 5 species R. zosimovichi , R. sidorenkoi , R. iconica ) in Rovno amber, plus "Bryocrypta " girafa Meunier (1904) and "B." capitosa Meunier (1904) in Baltic amber .
Sinallomyia [ 79]
Nom. nov
Valid
Zhang
Early Cretaceous
Yixian Formation
China
An archisargid brachyceran ; a replacement name for Allomyia Ren (1998).
Sinocretomyia [ 90]
Gen. et sp. nov
Valid
Zhang
Early Cretaceous
Laiyang Formation
China
An athericid . The type species is Sinocretomyia minuscula .
Stilobezzia pikei [ 100]
Sp. nov
Valid
Borkent
Cretaceous
Canadian amber
Canada
A species of Stilobezzia .
Stomatosema iljieteugeniae [ 77]
Sp. nov
Valid
Fedotova & Perkovsky
Late Eocene
Rovno amber
Ukraine
A cecidomyiid fly
Tipula paleopannonia [ 102]
Sp. nov
Valid
Engel & Gross
Miocene (ca. 11.3 Ma)
Paldau Formation
Austria
A crane fly , a species of Tipula .
Ephemeroptera
Hemiptera
Name
Novelty
Status
Authors
Age
Unit
Location
Notes
Images
Acaricoris robertae [ 107]
Sp. nov
Valid
Heiss & Poinar
Miocene
Dominican Republic
An aradid found in Dominican amber , a species of Acaricoris .
Arctorthezia baltica [ 108]
Sp. nov
Valid
Vea & Grimaldi
Eocene
Europe
An ortheziid found in Eocene Baltic amber , a species of Arctorthezia .
Breviscutum [ 109]
Gen. et sp. nov
Valid[ 110]
Yao et al.
Cretaceous
China
A primipentatomid pentatomoid . The type species is Breviscutum lunatum .
Burmacicada [ 111]
Gen. et sp. nov
Valid
Poinar & Kritsky
Cretaceous
Burmese amber
Myanmar
A cicada . The type species is Burmacicada protera .
Burmorthezia [ 108]
Gen et 2 sp. nov
Valid
Vea & Grimaldi
Cretaceous (98 Ma)
Burmese amber
Myanmar
An ortheziid . Genus contains two species: Burmorthezia kotejai and B. insolita .
Carventus bechlyi [ 107]
Sp. nov
Valid
Heiss & Poinar
Miocene
Dominican Republic
An aradid found in Dominican amber , a species of Carventus .
Cicadocoris varians [ 112]
Sp. nov
Disputed
Dong, Yao & Ren
Middle Jurassic
Jiulongshan Formation
China
A moss bug belonging to the family Progonocimicidae . Jiang & Huang (2017) considered this species to be a probable junior synonym of Cicadocoris sinensis Hong (1983).[ 113]
Cinemala [ 114]
Gen. et sp. nov
Valid
Shcherbakov
Middle or Late Jurassic
Bakhar Group
Mongolia
A member of Cicadomorpha belonging to the family Hylicellidae . The type species is C. contrasta .
Cintux [ 115]
Gen. et sp. nov
Valid
Stroiński & Szwedo
Palaeocene
France
A lophopid planthopper . The type species is Cintux menatensis .
Conjucella [ 114]
Gen. et sp. nov
Valid
Shcherbakov
Early Cretaceous
Zaza Formation
Russia
A member of Cicadomorpha belonging to the family Hylicellidae . The type species is C. selene .
Dellashara [ 116]
Gen. et sp. nov
Valid
Shcherbakov
Late Jurassic
Shar Teg Sequence
Mongolia
A member of Cicadomorpha belonging to the superfamily Membracoidea and the family Archijassidae . The type species is D. tega .
Dominicicada [ 111]
Gen. et sp. nov
Valid
Poinar & Kritsky
Dominican Republic
A cicada found in Dominican amber . The type species is Dominicicada youngi .
Edessa protera [ 117]
Sp. nov
Valid
Poinar & Thomas
Tertiary
Mexico
A pentatomid stink bug found in Mexican amber, a species of Edessa .
Hispanocader [ 118]
Gen. et sp. nov
Valid
Golub, Popov & Arillo
Albian
Escucha Formation
Spain
A relative of tingids . The type species is Hispanocader lisae .
Hoffeinsoria [ 119]
Gen. et sp. nov
Valid
Herczek & Popov
Eocene
Prussian Formation
Europe
An isometopine mirid found in Eocene Baltic amber . The type species is Hoffeinsoria robusta .
Jiania [ 120]
Gen. et 2 sp. nov
Valid
Wang & Szwedo in Wang, Szwedo & Zhang
Middle Jurassic
Daohugou Beds
China
A sinoalid froghopper . Genus contains two species: Jiania crebra and Jiania gracila . Fu, Cai & Huang (2018) considered J. gracila to be a junior synonym of J crebra .[ 121]
Kachinocoris [ 122]
Gen. et sp. nov
Valid
Heiss
Cretaceous
Burmese amber
Myanmar
A flat bug found in Cretaceous Burmese Amber. The type species is Kachinocoris brevipennis .
Kemobius [ 116]
Gen. et sp. nov
Valid
Shcherbakov
Middle Jurassic
Itat Formation
Russia
A member of Cicadomorpha belonging to the superfamily Membracoidea and the family Archijassidae . The type species is K. lux .
Kisa [ 116]
Gen. et sp. nov
Valid
Shcherbakov
Middle Jurassic
Itat Formation
Russia
A member of Cicadomorpha belonging to the superfamily Membracoidea and the family Archijassidae . The type species is K. fasciata .
Kubecola [ 116]
Gen. et sp. nov
Valid
Shcherbakov
Middle Jurassic
Itat Formation
Russia
A member of Cicadomorpha belonging to the superfamily Membracoidea and the family Archijassidae . The type species is K. guttatus .
Microaradus [ 123]
Gen. et sp. nov
Valid
Heiss & Poinar
Mesozoic
Burmese amber
Myanmar
An archaearadine aradid found in Mesozoic Burmese amber. The type species is Microaradus anticus .
Minyscapheus [ 124]
Gen. et sp. nov.
Valid
Poinar, Kritsky & Brown
Dominican Republic
A cicada found in Dominican amber . The type species is Minyscapheus dominicanus .
Mixorthezia dominicana [ 108]
Sp. nov
Valid
Vea & Grimaldi
Miocene
Dominican Republic
An ortheziid found in Miocene Dominican amber , a species of Mixorthezia .
Mixorthezia kozari [ 108]
Sp. nov
Valid
Vea & Grimaldi
Miocene
Dominican Republic
An ortheziid found in Miocene Dominican amber , a species of Mixorthezia .
Mongolocoris [ 125]
Gen. et comb. nov
Valid
Ryzhkova
Early Cretaceous
Mongolia
An enicocorid leptopodomorph heteropteran , a new genus for "Enicocoris " tibialis Popov, 1986.
Myanmezira [ 123]
Gen. et sp. nov
Valid
Heiss & Poinar
Mesozoic
Burmese amber
Myanmar
A mezirine aradid found in Mesozoic Burmese amber. The type species is Myanmezira longicornis .
Nesoproxius latocanus [ 107]
Sp. nov
Valid
Heiss & Poinar
Miocene
Dominican Republic
An aradid found in Dominican amber , a species of Nesoproxius .
Notonecta vetula [ 126]
sp nov
Valid
Zhang, Yao & Ren
Late Jurassic
Chijinqiao Formation
China
A notonectid ,a species of Notonecta .
Oropentatoma [ 109]
Gen. et sp. nov
Valid[ 127]
Yao et al.
Cretaceous
China
A primipentatomid pentatomoid . The type species is Oropentatoma epichara .
Poljanka hirsuta [ 128]
sp nov
Valid
Yang, Yao & Ren
Middle Jurassic
Jiulongshan Formation
China
A member of Sternorrhyncha , a protopsyllidiid ; a species of Poljanka .
Primipentatoma [ 109]
Gen. et 2 sp. nov
Valid[ 129]
Yao et al.
Cretaceous
China
A primipentatomid pentatomoid . Genus contains two species: Primipentatoma peregrina and Primipentatoma fangi .
Proneoproxius [ 107]
Gen. et sp. nov
Valid
Heiss & Poinar
Miocene
Dominican Republic
An aradid found in Dominican amber . The type species is Proneoproxius cornutus .
Purbecellus [ 114]
Gen. et comb. nov
Valid
Shcherbakov
Early Cretaceous
Purbeck Group
United Kingdom
A member of Cicadomorpha belonging to the superfamily Membracoidea and the family Archijassidae . The type species is "Cicadellium " psocus Westwood (1854).
Quadrocoris [ 109]
Gen. et sp. nov
Valid[ 130]
Yao et al.
Cretaceous
China
A primipentatomid pentatomoid . The type species is Quadrocoris radius .
Sinaldocader rasnitsyni [ 131]
Sp. nov
Valid
Golub & Popov
Turonian
Kazakhstan
A lace bug . Originally described as a species of Sinaldocader ; Popov & Golub (2019) transferred this species to the separate genus Kzylcader .[ 132]
Sinoala [ 120]
Gen. et sp. nov
Valid
Wang & Szwedo in Wang, Szwedo & Zhang
Middle Jurassic
Daohugou Beds
China
A sinoalid froghopper . The type species is Sinoala parallelivena .
Sunotettigarcta hirsuta [ 133]
sp nov
Valid
Li, Wang & Ren in Li et al.
Middle Jurassic
Daohugou Beds
China
A tettigarctid , a species of Sunotettigarcta .
Ulanocoris [ 125]
Gen. et 2 sp. nov
Valid
Ryzhkova
Early Cretaceous
Mongolia
An enicocorid leptopodomorph heteropteran . Genus contains two species: Ulanocoris femoralis and U. grandis .
Venustsalda [ 134]
Gen. et sp. nov
Valid
Zhang et al.
Lower Cretaceous
Yixian Formation
China
A saldid heteropteran . The type species is Venustsalda locella .
Hymenoptera
Name
Novelty
Status
Authors
Age
Unit
Location
Notes
Images
Agapia [ 135]
Gen. et sp. nov
Valid
Kopylov
Late Cretaceous
Russia
An ichneumonid . The type species is Agapia sukatchevae .
Agapteron [ 135]
Gen. et sp. nov
Valid
Kopylov
Late Cretaceous
Russia
An ichneumonid . The type species is Agapteron popovi .
Archimyrmex wedmannae [ 136]
Sp. nov
Valid
Dlussky
Middle Eocene
Messel Pit
Germany
A myrmeciine ant .
Archimyrmex wedmannae
Anthophorula (Anthophorula) persephone [ 137]
Sp. nov
Valid
Engel in Engel et al.
Early Miocene
Dominican Republic
An exomalopsine apid bee found in Dominican amber , a species of Anthophorula .
Atefia [ 138]
Gen. et sp. nov
Valid[ 139]
Krogmann et al.
Early Cretaceous
Crato Formation
Brazil
A tenthredinoid hymenopteran sharing morphological characters with the families Pergidae and Diprionidae . The type species is Atefia rasnitsyni .
Boltonidris [ 140]
Gen. et sp. nov
Valid
Radchenko & Dlussky
Late Eocene
Rovno amber
Ukraine
A stenammine myrmicine ant found in Rovno amber. The type species is Boltonidris mirabilis .
Bombus (Bombus) randeckensis [ 141]
Sp. nov
Valid
Wappler & Engel in Wappler et al.
Miocene
Germany
A bumblebee .
Casaleia eocenica [ 142] [ 143]
Sp nov
valid
Dlussky & Wedmann
Lutetian
Messel Formation
Germany
An amblyoponine ant.
Casaleia eocenica
Cascoscelio [ 144]
Gen. et sp. nov.
Valid
Poinar in Poinar & Buckley
Early Cretaceous
Burmese amber
Myanmar
A platygastrid wasp, type species is C. incassus .
Cascoscelio incassa
Cephalopone [ 142]
Gen. et 2 sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A poneromorph ant . Two species: C. potens & C. grandis .
Cephalopone grandis
Cratoenigma [ 145]
Gen. et sp. nov
Valid
Krogmann & Nél
Early Cretaceous (Aptian )
Crato Formation
Brazil
A probable relative of xiphydriids and euhymenopterans . The type species is C. articulata .
Cretevania soplaensis [ 146]
Sp. nov
Valid
De La Fuente, Peñalver & Ortega-Blanco
Albian
Spain
An evaniid , a species of Cretevania .
Cretorhyssalus [ 147]
Gen. et sp. nov
Valid
Belokobylskij
Cenomanian
Ola Formation
Russia
A braconid wasp . Type species is Cretorhyssalus brevis .
Cyrtopone [ 142]
Gen. et 4 sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel Formation
Germany
A ponerinae ant . Genus contains four species: Cyrtopone curiosa , C. elongata , C. microcephala and C. striata .
C. striata
Gulgonga [ 148]
Gen. et sp. nov
Valid
Oberprieler, Rasnitsyn & Brothers
Late Jurassic
Talbragar Fish Bed
Australia
A praeaulacid evanioid . The type species is Gulgonga beattiei .
Haidomyrmex scimitarus [ 149]
Sp. nov
Valid
Barden & Grimaldi
Late Aptian to Early Cenomanian
Burmese amber
Myanmar
A Sphecomyrmin ant
Haidomyrmex scimitarus
Haidomyrmex zigrasi [ 149]
Sp. nov
Valid
Barden & Grimaldi
Campanian
Burmese amber
Myanmar
A Sphecomyrmin ant
Haidomyrmex zigrasi
Hyptia deansi [ 150]
Sp. nov
Valid
Jennings, Krogmann & Mew
Late Oligocene or early Miocene
Mexico
An evaniid hymenopteran , a species of Hyptia .
Ibalia (Ibalia) electra [ 151]
Sp. nov
Valid
Engel & Liu
Eocene
Europe
An ibaliid found in Baltic amber , a species of Ibalia .
Lancepyris [ 152]
Gen. et sp. nov
Valid
Azevedo & Azar
Early Cretaceous
Lebanon
A lancepyrine bethylid found in Early Cretaceous Lebanese amber. The type species is Lancepyris opertus .
Lonchodryinus balticus [ 153]
Sp. nov
Valid
Olmi & Guglielmino
Late Eocene
Europe
A dryinid found in Baltic amber , a species of Lonchodryinus .
Magadanobracon [ 147]
Gen. et 2 sp. nov
Valid
Belokobylskij
Cenomanian
Ola Formation
Russia
A braconid wasp. The type species is M. rasnitsyni ; a second species is M. zherikhini .
Messelepone [ 142]
Gen. et sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A poneromorph ant . The type species is M. leptogenoides .
Messelepone leptogenoides
Mirabythus [ 154]
Gen. et 2 sp. nov
Valid
Cai, Shih & Ren in Cai et al.
Early Cretaceous
Yixian Formation
China
A scolebythid chrysidoid . The genus contains two species: Mirabythus lechrius and M. liae .
Pachycondyla eocenica [ 142]
Sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A Ponerinae ant .
Pachycondyla eocenica
Pachycondyla lutzi [ 142]
Sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A ponerine ant
Pachycondyla lutzi
Pachycondyla? messeliana [ 142]
Sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A ponerine ant .
Pachycondyla? messeliana
Pachycondyla minuta [ 142] [ 155]
Sp. nov
synonym
Dlussky & Wedmann
Eocene
Messel pit
Germany
A ponerine ant , name occupied, renamed P. parvula .
Pachycondyla parvula
Pachycondyla petiolosa [ 142]
Sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A ponerine ant .
Pachycondyla petiolosa
Pachycondyla petrosa [ 142]
Sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A poneromorph ant , a species of Pachycondyla .
Pachycondyla petrosa
Platyxyela [ 156]
Gen. et sp. nov
Valid
Wang, Shih & Ren
Middle Jurassic
Jiulongshan Formation
China
A macroxyeline xyelid sawfly . The type species is Platyxyela unica .
Praeichneumon dzhidensis [ 157]
sp nov
Valid
Kopylov
Lower Cretaceous
Russia
A praeichneumonid ichneumonoid , a species of Praeichneumon .
Praeichneumon khamardabanicus [ 157]
sp nov
Valid
Kopylov
Lower Cretaceous
Russia
A praeichneumonid ichneumonoid, a species of Praeichneumon .
Praeichneumon zakhaaminicus [ 157]
sp nov
Valid
Kopylov
Lower Cretaceous
Russia
A praeichneumonid ichneumonoid, a species of Praeichneumon .
Prionomyrmex wappleri [ 136]
Sp. nov
Valid
Dlussky
Late Oligocene
Germany
A myrmeciine ant .
Prionomyrmex wappleri
Protodryinus [ 153]
Gen. et sp. nov
Valid
Olmi & Guglielmino
Late Eocene
Europe
A dryinid found in Baltic amber . The type species is Protodryinus eocenicus .
Protopone? dubia [ 142]
Sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel Formation
Germany
A ponerine ant
Protopone? dubia
Protopone germanica [ 142]
Sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A ponerine ant
Protopone germanica
Protopone magna [ 142]
Sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A ponerine ant
Protopone magna
Protopone oculata [ 142]
Sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A ponerine ant
Protopone oculata
Protopone sepulta [ 142]
Sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A ponerine ant
Protopone sepulta
Protopone vetula [ 142]
Sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
A ponerine ant
Protopone vetula
Pseudectatomma [ 142]
Gen. et 2 sp. nov
Valid
Dlussky & Wedmann
Eocene
Messel pit
Germany
An Ectatommin ant . Two species: P. eocenica and P. striatula .
Pseudectatomma eocenica
Trichelyon [ 158]
Gen. et sp. nov
Disputed
Ortega-Blanco & Engel in Ortega-Blanco, Singh & Engel
Early Eocene (Ypresian )
India
A rhysipoline exothecine braconid wasp found in Cambay amber. The type species is Trichelyon tadkeshwarense . Belokobylskij & Manukyan (2024) considered Trichelyon to be a junior synonym of the genus Clinocentrus , though the authors maintained T. tadkeshwarense as a distinct species within the latter genus, resulting in a new combination Clinocentrus tadkeshwarense .[ 159]
Urotryphon baikurensis [ 135]
Sp. nov
Valid
Kopylov
Late Cretaceous
Russia
An ichneumonid , a species of Urotryphon .
Zatania electra [ 160]
Sp. nov
Valid
Lapolla, Kallal & Brady
Eocene or Miocene
Dominican amber
Dominican Republic
A formicine ant , a species of Zatania .
Zatania electra
Mecoptera
Name
Novelty
Status
Authors
Age
Unit
Location
Notes
Images
Baltipanorpa [ 161]
Gen. et sp. nov
Valid
Krzemiński & Soszyńska-Maj
Middle Eocene
Baltic amber
Poland
A panorpid scorpionfly . The type species is B. damzeni .
Exilibittacus [ 162]
Gen. et sp. nov
Valid
Yang, Ren & Shih
Middle Jurassic
Jiulongshan Formation
China
A bittacid hangingfly . The type species is E. lii .
Exilibittacus lii
Jeholopsyche bella [ 163]
Sp. nov
Valid
Qiao, Shih & Ren
Early Cretaceous
Yixian Formation
China
An aneuretopsychid mecopteran .
Jeholopsyche completa [ 163]
Sp. nov
Valid
Qiao, Shih & Ren
Early Cretaceous
Yixian Formation
China
An aneuretopsychid mecopteran .
Jeholopsyche maxima [ 163]
Sp. nov
Valid
Qiao, Shih & Ren
Early Cretaceous
Yixian Formation
China
An aneuretopsychid mecopteran .
Juracimbrophlebia [ 164]
Gen. et sp. nov
Valid
Wang, Labandeira, Shih & Ren in Wang et al.
Middle Jurassic (Bathonian or Callovian)
Jiulongshan Formation
China
A cimbrophlebiid mecopteran . Adults with Yimaia ginkglaean leaf mimicry . The type species is J. ginkgofolia .
Juracimbrophlebia ginkgofolia
Megabittacus spatiosus [ 162]
Sp. nov
Valid
Yang, Ren & Shih
Late Jurassic or Early Cretaceous
Yixian Formation
China
A hangingfly , a species of Megabittacus .
Orthophlebia nervulosa [ 165]
sp nov
Valid
Qiao, Shih & Ren
Middle Jurassic
Jiulongshan Formation
China
A mecopteran , a species of Orthophlebia .
Orthophlebia riccardii [ 166]
Sp. nov
Valid
Petrulevičius & Ren
Middle Jurassic
Jiulongshan Formation
China
A mecopteran . Originally described as a species of Orthophlebia , subsequently transferred to the separate genus Gigaphlebia .[ 167]
Orthophlebia stigmosa [ 165]
sp nov
Valid
Qiao, Shih & Ren
Middle Jurassic
Jiulongshan Formation
China
A mecopteran . Originally described as a species of Orthophlebia , subsequently transferred to the separate genus Longiphlebia .[ 167]
Perfecticimbrophlebia [ 168]
Gen. et sp. nov
Valid
Yang, Shih & Ren in Yang et al.
Middle Jurassic
Jiulongshan Formation
China
A cimbrophlebiid mecopteran . The type species is Perfecticimbrophlebia laetus .
Preanabittacus validus [ 168]
sp nov
Valid
Yang, Shih & Ren in Yang et al.
Middle Jurassic
Jiulongshan Formation
China
A hangingfly , a species of Preanabittacus .
Neuroptera
Name
Novelty
Status
Authors
Age
Unit
Location
Notes
Images
Alloepipsychopsis [ 169]
Gen. et sp. nov
Valid
Makarkin, Yang, et al.
Barremian
Yixian Formation
China
A psychopsid . Type species is Alloepipsychopsis lata .
Chorilingia [ 170]
Gen. et 4 sp. nov
Valid
Shi et al.
Middle Jurassic
Jiulongshan Formation
China
A grammolingiid neuropteran . Genus contains four species: of C. euryptera , C. parvica , C. translucida and C. peregrina .
Guyiling [ 171]
Gen. et sp. nov
Valid
Shi et al.
Early Cretaceous
Yixian Formation
China
A member of Myrmeleontiformia of uncertain phylogenetic placement. The type species is G. jianboni .
Hallucinochrysa [ 172]
Gen. et sp. nov
Valid
Pérez-de la Fuente, Delclòs, Peñalver & Engel in Pérez-de la Fuente et al.
Early Cretaceous (Albian)
Spain
A chrysopoid neuropteran (a relative of the green lacewings ). The type species is Hallucinochrysa diogenesi .
Kareninoides [ 173]
Gen. et sp. et comb. nov
Valid
Yang, Makarkin & Ren
Early Cretaceous
La Huerguina Formation
Yixian Formation
China
Spain
A mesochrysopid neuropteran . The type species is Kareninoides lii ; genus might also contain "Armandochrysopa" inexpecta Nel et al. (2005).
Leptolingia oblonga [ 174]
Sp. nov
Valid
Khramov
Late Jurassic
Mongolia
A grammolingiid neuropteran , a species of Leptolingia .
Litholingia longa [ 174]
Sp. nov
Valid
Khramov
Early or Middle Jurassic
Kyrgyzstan
A grammolingiid neuropteran , a species of Litholingia .
Parakseneura [ 175]
Gen. et 11 sp. nov
Valid
Yang, Makarkin & Ren in Yang et al.
Middle Jurassic
Jiulongshan Formation
China
A parakseneurid neuropteran . Genus contains 11 species.
Protoaristenymphes daohugouensis [ 173]
Sp. nov
Valid
Yang, Makarkin & Ren
Middle Jurassic
Daohugou Beds
China
A mesochrysopid neuropteran , a species of Protoaristenymphes .
Protolingia [ 174]
Gen. et sp. nov
Valid
Khramov
Early or Middle Jurassic
Kyrgyzstan
A grammolingiid neuropteran . The type species is Protolingia mira .
Pseudorapisma [ 175]
Gen. et 3 sp. nov
Valid
Yang, Makarkin & Ren in Yang et al.
Middle Jurassic
Jiulongshan Formation
China
A parakseneurid neuropteran . Genus contains 3 species: P. jurassicum , P. maculatum and P. angustipenne .
Shuraboneura [ 175]
Gen. et sp. nov
Valid
Khramov & Makarkin in Yang et al.
Early or Middle Jurassic
Sogul Formation
Kyrgyzstan
A parakseneurid neuropteran . The type species is Shuraboneura ovata .
Notoptera
Odonatoptera
Name
Novelty
Status
Authors
Age
Unit
Location
Notes
Images
Austroprotolindenia [ 182]
Gen. et sp. nov
Valid
Beattie & Nel
Late Jurassic
Talbragar Fossil Fish Bed
Australia
A dragonfly . The type species is Austroprotolindenia jurassica .
Azaroneura [ 183]
Gen. et sp. nov
Valid
Nel et al.
Permian (Kungurian)
Koshelevka Formation
Russia
A protozygopteran . The type species is A. permiana .
Balticoagrion [ 184]
Gen. et sp. nov
Valid
Bechly
Eocene
Europe
A damselfly found in Baltic amber . The type species is Balticoagrion paulyi .
Decoraeshna [ 185]
Gen. et sp. nov
Valid
Li et al.
Early Cretaceous
Yixian Formation
China
A progobiaeshnid dragonfly . The type species is Decoraeshna preciosus .
Engellestes [ 183]
Gen. et sp. nov
Valid
Nel et al.
Permian (Kungurian)
Koshelevka Formation
Russia
A protozygopteran . The type species is E. chekardensis .
Epilestes angustapterix [ 183]
Sp. nov
Valid
Nel et al.
Late Permian
Russia
A protozygopteran.
Germanostenolestes [ 186]
Gen. et sp. nov
Valid
Nel & Fleck
Middle Miocene
Germany
A sieblosiid epiproctophoran odonate . The type species is Germanostenolestes lutzi .
Karatawia sinica [ 187]
Sp. nov
Valid
Li, Nel & Ren in Li et al.
Middle Jurassic
Jiulongshan Formation
China
A campterophlebiid odonate ,a species of Karatawia .
Kennedya azari [ 183]
Sp. nov
Valid
Nel et al.
Permian (Kungurian)
Koshelevka Formation
Russia
A protozygopteran.
Kennedya ivensis [ 183]
Sp. nov
Valid
Nel et al.
Late Permian
Russia
A protozygopteran.
Kennedya pritykinae [ 183]
Sp. nov
Valid
Nel et al.
Late Permian
Russia
A protozygopteran.
Luiseia [ 183]
Gen. et sp. nov
Valid
Nel et al.
Carboniferous/Permian boundary
Bursum Formation
United States
A protozygopteran. The type species is L. breviata .
Mongoliaeshna exiguusens [ 185]
Sp. nov
Valid
Li et al.
Early Cretaceous
Yixian Formation
China
A progobiaeshnid dragonfly , a species of Mongoliaeshna .
Mongoliaeshna hadrens [ 185]
Sp. nov
Valid
Li et al.
Early Cretaceous
Yixian Formation
China
A progobiaeshnid dragonfly , a species of Mongoliaeshna .
Oligomazon [ 188]
Gen. et comb. nov
Valid
Zessin & Brauckmann
Carboniferous (Pennsylvanian )
United States
A member of Meganisoptera , a paralogid ; a new genus for "Oligotypus " makowskii Carpenter & Richardson (1971).
Paragilsonia [ 189]
Gen. et sp. nov (but see notes)
Valid
Zhang, Hong & Su in Su et al.
Carboniferous (Namurian )
China
A member of Odonatoptera ; the type species is Paragilsonia orientalis . Li et al. (2013) synonymized Paragilsonia with the genus Tupus Sellards (1906); however, they retained P. orientalis as a distinct species within the latter genus.[ 190]
Paralogobora [ 188]
Gen. et sp. nov
Valid
Zessin & Brauckmann
Early Permian
Czech Republic
A member of Odonata . The type species is Paralogobora guentherpetersi .
Permolestes sheimogorai [ 183]
Sp. nov
Valid
Nel et al.
Late Permian
Russia
A protozygopteran.
Permolestes soyanaiensis [ 183]
Sp. nov
Valid
Nel et al.
Late Permian
Russia
A protozygopteran.
Progoneura grimaldii [ 183]
Sp. nov
Valid
Nel et al.
Early Permian
Wellington Formation
United States
A protozygopteran.
Pseudocymatophlebia boda [ 191]
Sp. nov
Valid
Li, Nel & Ren in Li et al.
Early Cretaceous
China
An aktassiid odonate , a species of Pseudocymatophlebia .
Sinierasiptera [ 189]
Gen. et sp. nov (but see notes)
Valid
Zhang, Hong & Su in Su et al.
Carboniferous (Namurian )
China
A member of Odonatoptera ; the type species is Sinierasiptera jini . Li et al. (2013) synonymized Sinierasiptera with the genus Erasipterella Brauckmann (1983); however, they retained S. jini as a distinct species within the latter genus.[ 190]
Sinojagoria cancellosa [ 185]
Sp. nov
Valid
Li et al.
Early Cretaceous
Yixian Formation
China
A gomphaeschnid dragonfly , a species of Sinojagoria .
Sinojagoria magna [ 185]
Sp. nov
Valid
Li et al.
Early Cretaceous
Yixian Formation
China
A gomphaeschnid dragonfly , a species of Sinojagoria .
Solikamptilon pectinatus [ 183]
Sp. nov
Valid
Nel et al.
Late Permian
Russia
A protozygopteran.
Orthopterida
Raphidioptera
Other
Name
Novelty
Status
Authors
Age
Unit
Location
Notes
Images
Anebos [ 202]
Gen. et sp. nov
Valid
Garwood et al.
Late Pennsylvanian
Assise de Montceau Formation
France
A polyneopteran insect of uncertain affinities. The type species is Anebos phrixos .
Anebos phrixos
Bechala [ 203]
Gen. et sp. nov
Valid
Ilger & Brauckmann
Late Carboniferous
Germany
A member of Odonatoptera , assigned to the new family Bechalidae . Originally interpreted as a megasecopteran .[ 204] Type species is Bechala sommeri .
Bruayaphis [ 205]
Gen. et sp. nov
Valid
Nel, Szwedo & Roques in Nel et al.
Late Carboniferous
France
A member of Paraneoptera of uncertain phylogenetic placement. The type species is Bruayaphis oudardi .
Cucullistriga [ 206]
Gen. et sp. nov
Valid
Aristov & Rasnitsyn
Kungurian
Russia
A member of Grylloblattida/Eoblattida belonging to the family Idelinellidae ; a relative of ice crawlers or cockroaches .[ 207] Type species is Cucullistriga cucullata .
Darekia sanguinea [ 208]
Gen. et sp. nov
Valid
Prokop et al.
Upper Carboniferous
Upper Silesian Coal Basin
Poland
A stem winged insect.
Eochauliodes [ 209]
Gen. et sp. nov
Valid
Liu et al.
Middle Jurassic (Bathonian-Callovian)
Jiulongshan Formation
China
A fishfly . The type species is Eochauliodes striolatus .
Eospilopsyllus [ 210]
Gen. et sp. nov
Valid
Beaucournu & Perrichot in Perrichot, Beaucournu & Velten
Early Miocene
Dominican Republic
A pulicid flea found in Dominican amber . The type species is Eospilopsyllus kobberti .
Geinitzia aristovi [ 211]
sp nov
Valid
Cui, Storozhenko & Ren
Middle Jurassic
Jiulongshan Formation
China
A geinitziid , a member of Polyneoptera /Gryllones belonging to the group Reculida ;[ 207] a species of Geinitzia .
Gymnopollisthrips [ 212]
Gen. et 2 sp. nov.
Valid
Peñalver, Nel & Nel in Peñalver et al.
Early Cretaceous
Spain
A melanthripid thrip . Genus contains two species: Gymnopollisthrips minor and G. maior .
Liadotaulius daohugouensis [ 213]
sp nov
Valid
Wu & Huang
Middle Jurassic
Daohugou Beds
China
A caddisfly , a species of Liadotaulius .
Netoxena [ 214]
Nom. nov
Valid
Sohn in Sohn et al.
Early Cretaceous (late Aptian)
Crato Formation
Brazil
A lepidopteran , possibly a member of Eolepidopterigidae ; a replacement name for Xena Martins-Neto (1999).
Pseudopulex [ 215]
Gen. et 2 sp. nov
Valid
Gao, Shih, & Ren in Gao et al.
Middle Jurassic to Early Cretaceous
China
A flea-like insect. The genus contains 2 species: P. jurassicus and P. magnus .
Rasstriga [ 206]
Gen. et sp. nov
Valid
Aristov & Rasnitsyn
Upper Carboniferous (Desmoinesian)
United States
A member of Grylloblattida/Eoblattida probably belonging to the family Idelinellidae ; a relative of ice crawlers or cockroaches .[ 207] Its type species is Rasstriga americana .
Scutistriga [ 206]
Gen. et sp. nov
Valid
Aristov & Rasnitsyn
Kungurian
Russia
A member of Grylloblattida/Eoblattida belonging to the family Idelinellidae ; a relative of ice crawlers or cockroaches .[ 207] Its type species is Scutistriga scutata .
Sharasialis [ 216]
Gen. et sp. nov
Valid
Ponomarenko
Late Jurassic
Mongolia
An alderfly . The type species is Sharasialis fusiformis .
Sinoprotodiplatys [ 217]
Gen. et sp. nov
Valid
Nel et al.
Early Cretaceous
Yixian Formation
China
An earwig belonging to the family Protodiplatyidae . The type species is Sinoprotodiplatys zhangi .
Strigulla [ 206]
Gen. et comb. nov
Valid
Aristov & Rasnitsyn
Kungurian
Russia
A member of Grylloblattida/Eoblattida belonging to the family Idelinellidae ; a relative of ice crawlers or cockroaches .[ 207] A new genus for "Euryptilon " cuculiophoris (Aristov, 2002).
Strudiella [ 218]
Gen. et sp. nov.
Valid
Garrouste et al.
Late Devonian
Belgium
The first complete Late Devonian insect, probably terrestrial. The type species is Strudiella devonica .
Zdenekia silesiensis [ 208]
Species
Valid
Prokop et al.
Upper Carboniferous
Upper Silesian Coal Basin
Poland
A stem winged insect.
General research
References
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^ Yongjie Wang; Conrad C. Labandeira; Chungkun Shih; Qiaoling Ding; Chen Wang; Yunyun Zhao; Dong Ren (2012). "Jurassic mimicry between a hangingfly and a ginkgo from China" . Proceedings of the National Academy of Sciences of the United States of America . 109 (50): 20514– 20519. Bibcode :2012PNAS..10920514W . doi :10.1073/pnas.1205517109 . PMC 3528590 . PMID 23184994 .
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^ Julián F. Petrulevičius; Dong Ren (2012). "A new species of "Orthophlebiidae" (Insecta: Mecoptera) from the Middle Jurassic of Inner Mongolia, China" (PDF) . Revue de Paléobiologie, Genève . spécial 11: 311– 315.
^ a b Agnieszka Soszyńska-Maj; Wiesław Krzemiński; Katarzyna Kopeć; Yizi Cao; Dong Ren (2018). "Large Jurassic scorpionflies belonging to a new subfamily of the family Orthophlebiidae (Mecoptera)". Annales Zoologici . 68 (1): 85– 92. doi :10.3161/00034541ANZ2018.68.1.004 . S2CID 90298406 .
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^ Vladimir N. Makarkin; Qiang Yang; YuanYuan Peng; Dong Ren (2012). "A comparative overview of the neuropteran assemblage of the Lower Cretaceous Yixian Formation (China), with description of a new genus of Psychopsidae (Insecta: Neuroptera)". Cretaceous Research . 35 : 57– 68. Bibcode :2012CrRes..35...57M . doi :10.1016/j.cretres.2011.11.013 .
^ Chao-Fan Shi; Yong-Jie Wang; Qiang Yang; Dong Ren (2012). "Chorilingia (Neuroptera: Grammolingiidae): a new genus of lacewings with four species from the Middle Jurassic of Inner Mongolia, China". Alcheringa: An Australasian Journal of Palaeontology . 36 (3): 309– 318. Bibcode :2012Alch...36..309S . doi :10.1080/03115518.2012.644994 . S2CID 84255854 .
^ Chaofan Shi; Olivier Béthoux; Chungkun Shih; Dong Ren (2012). "Guyiling jianboni gen. et sp.n., an antlion-like lacewing, illuminating homologies and transformations in Neuroptera wing venation". Systematic Entomology . 37 (3): 617– 631. Bibcode :2012SysEn..37..617S . doi :10.1111/j.1365-3113.2012.00633.x . S2CID 84467796 .
^ Ricardo Pérez-de la Fuente; Xavier Delclòs; Enrique Peñalver; Mariela Speranza; Jacek Wierzchos; Carmen Ascaso; Michael S. Engel (2012). "Early evolution and ecology of camouflage in insects" . Proceedings of the National Academy of Sciences of the United States of America . 109 (52): 21414– 21419. Bibcode :2012PNAS..10921414P . doi :10.1073/pnas.1213775110 . PMC 3535654 . PMID 23236135 .
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^ Yingying Cui; Olivier Béthoux; Nan Yang; Dong Ren (2023). "New data and systematic considerations on the Blattogryllopterida (Insecta: Grylloblattida) from the Jurassic Daohugou locality in China". Palaeoentomology . 6 (2): 124– 132. doi :10.11646/palaeoentomology.6.2.4 . S2CID 258399002 .
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^ Yingying Cui; Stephan Brauner; Joerg W. Schneider; Olivier Béthoux (2021). "Grylloblattidan insects from Sperbersbach and Cabarz (Germany), two new early Permian and insect-rich localities" . Journal of Paleontology . 96 (2): 355– 374. doi :10.1017/jpa.2021.101 . S2CID 243486826 .
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^ André Nel; Gunther Fleck (2012). "A new genus and species of Sieblosiidae from the Middle Miocene of Germany (Odonata: Epiproctophora)" (PDF) . Zootaxa . 3582 : 64– 68. doi :10.11646/zootaxa.3582.1.7 .
^ Yong-Jun Li; André Nel; Dong Ren; Bing-Lan Zhang; Hong Pang (2012). "Reassessment of the Jurassic damsel-dragonfly genus Karatawia (Odonata: Campterophlebiidae)" (PDF) . Zootaxa . 3417 : 64– 68. doi :10.11646/zootaxa.3417.1.4 .
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^ Yongjun Li; Gang Han; André Nel; Dong Ren; Hong Pang; Xueling Liu (2012). "A new fossil petalurid dragonfly (Odonata: Petaluroidea: Aktassiidae) from the Cretaceous of China". Alcheringa: An Australasian Journal of Palaeontology . 36 (3): 319– 322. Bibcode :2012Alch...36..319L . doi :10.1080/03115518.2012.644995 . S2CID 128394986 .
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^ Olivier Béthoux; Junjie Gu; Yanli Yue; Dong Ren (2012). "Miamia maimai n. sp., a new Pennsylvanian stem-orthopteran insect, and a case study on the application of cladotypic nomenclature" . Fossil Record . 15 (2): 103– 113. doi :10.1002/mmng.201200008 .
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