List of European dinosaurs
The Tethys Ocean splitting Laurasia from Gondwana .
Dinosaurs evolved partway through the Triassic period of the Mesozoic era, around 230 Ma (million years ago). At that time, the earth had one supercontinental landmass, called Pangaea , of which Europe was a part. So it remained throughout the Triassic. By the start of the Jurassic period, some 30 million years later, the supercontinent began to split into Laurasia and Gondwana . The largest inlet from Panthalassa , the superocean that surrounded Pangaea, was called the Tethys Ocean , and as this inlet cut deeper into the supercontinent, much of Europe was flooded.
By the Cretaceous , from 145 to 66 million years ago, the continents were beginning to approach their present shapes, but not their present positions, and Europe remained tropical. At times, it was a chain of island-microcontinents including Baltica and Iberia .
Europe is relatively rich in fossils from the Jurassic-Cretaceous boundary, and much of what is known about European dinosaurs dates from this time. During the Maastrichtian the end of the Cretaceous dinosaurs were dominating western and Central Europe as the Tremp Formation in Spain dates back to that age. Examples of dinosaurs from Maastrichtian Europe are Struthiosaurus and Canardia .
Criteria for inclusion
List of European dinosaurs
Valid genera
Name
Year
Formation
Location
Notes
Images
Abditosaurus
2022
Tremp Formation (Late Cretaceous , Maastrichtian )
Spain
Larger and distantly related to other European sauropods, suggesting it was an immigrant from another continent[ 1]
Acanthopholis
1867
Chalk Group (Late Cretaceous , Cenomanian )
England
Possessed keeled oval scutes as well as long spines
Adynomosaurus
2019
Tremp Formation (Late Cretaceous , Maastrichtian )
Spain
Relatively basal despite its late age[ 2]
Aepisaurus
1852
Grès verts hevétiques (Early Cretaceous , Albian )
France
Known only from a humerus which shares some features with camarasaurids and titanosaurs[ 3]
Ajkaceratops
2010
Csehbánya Formation (Late Cretaceous , Santonian )
Hungary
Originally thought to be a ceratopsian although this identification has been doubted by a later study[ 4]
Allosaurus
1877
Lourinhã Formation (Late Jurassic , Kimmeridgian )
Portugal
Three species, including the type, are known from the United States. The European species may be a synonym of A. fragilis [ 5]
Alocodon
1973
Cabaços Formation (Late Jurassic , Oxfordian )
Portugal
Had vertical grooves on its teeth
Altispinax
1923
Wadhurst Clay Formation (Early Cretaceous , Valanginian )
England
Possessed elongated neural spines that may have supported a hump-like structure as in Concavenator
Amanzia
2020
Reuchenette Formation (Late Jurassic , Kimmeridgian )
Switzerland
One bone preserves fossilized cartilage.[ 6] Originally believed to be a species of Ornithopsis or Cetiosauriscus
Ampelosaurus
1995
Argiles et Grès à Reptiles Formation , Grès de Labarre , Gres de Saint-Chinian , Marnes Rouges Inférieures Formation (Late Cretaceous , Maastrichtian )
France
Had three different types of osteoderms, including spines, plates and bulbs
Anoplosaurus
1879
Cambridge Greensand (Early Cretaceous , Albian )
England
Considered an ankylosaur despite the fact no osteoderms have been found. Historically suggested to be an ornithopod
Aragosaurus
1987
Villar del Arzobispo Formation (Early Cretaceous , Berriasian )
Spain
Reportedly had a unique ischium
Arcovenator
2014
Argiles et Grès à Reptiles Formation (Late Cretaceous , Campanian )
France
Most closely related to abelisaurids from India and Madagascar[ 7]
Arenysaurus
2009
Tremp Formation (Late Cretaceous , Maastrichtian )
Spain
One of the last lambeosaurines prior to their extinction
Aristosuchus
1887
Wealden Group (Early Cretaceous , Barremian )
England
May have had long claws based on referred unguals
Asylosaurus
2007
Avon Fissure Fill (Late Triassic , Rhaetian )
England
Remains originally identified as Thecodontosaurus
Atsinganosaurus
2010
Argiles et Grès à Reptiles Formation (Late Cretaceous , Maastrichtian )
France
Belongs to a unique clade of southwestern European titanosaurs[ 8]
Aviatyrannis
2003
Alcobaça Formation (Late Jurassic , Oxfordian to Tithonian )
Portugal
Although originally described as a tyrannosauroid, it may in fact be the oldest known ornithomimosaur[ 9]
Barilium
2010
Wadhurst Clay Formation (Early Cretaceous , Valanginian )
England
Robust with strong vertebrae and short neural spines
Baryonyx
1986
Weald Clay (Early Cretaceous , Barremian )
England
One specimen was found with the remains of fish and a juvenile iguanodont in its stomach, suggesting it was a generalist predator[ 10]
Betasuchus
1932
Maastricht Formation (Late Cretaceous , Maastrichtian )
Netherlands
The first terrestrial vertebrate named from the Maastrichtian stage[ 11]
Blasisaurus
2010
Arén Formation (Late Cretaceous , Maastrichtian )
Spain
Only known from a partial skull but can be distinguished from contemporary lambeosaurines[ 12]
Bothriospondylus
1875
Kimmeridge Clay (Late Jurassic , Kimmeridgian )
England
Several specimens have been assigned to this genus, but most of them have been reclassified into different genera
Bradycneme
1975
Sânpetru Formation (Late Cretaceous , Maastrichtian )
Romania
Potentially an alvarezsaurid[ 6]
Brighstoneus
2021
Wessex Formation (Early Cretaceous , Barremian )
England
Possessed a long snout tipped with a low bump
Burianosaurus
2017
Peruc-Korycany Formation (Late Cretaceous , Cenomanian )
Czech Republic
May be closely allied to the rhabdodontids[ 13]
Calamosaurus
1891
Wessex Formation (Early Cretaceous , Barremian )
England
Only known from two cervical vertebrae. Several bones have been referred to it over the years but there is no way to prove all of them belonged to the same genus
Calamospondylus
1866
Wessex Formation (Early Cretaceous , Hauterivian )
England
Only definitively known from a sacrum and associated pelvic elements
Caletodraco
2024
Chalk of the Pays de Caux (Late Cretaceous , Cenomanian )
France
Possibly the first known non-South American furileusaurian abelisaurid[ 14]
Callovosaurus
1980
Oxford Clay (Middle Jurassic , Callovian )
England
The oldest known dryosaurid[ 15]
Calvarius
2023
Tremp Formation (Late Cretaceous , Maastrichtian )
Spain
May have been a cursorial biped similar to basal ornithopods despite being more derived
Camarillasaurus
2014
Camarillas Formation (Early Cretaceous , Barremian )
Spain
Described as a ceratosaur[ 16] but has since been reinterpreted as a spinosaurid[ 17]
Camelotia
1985
Westbury Formation (Late Triassic to Early Jurassic , Rhaetian to Hettangian )
England
One of the largest known non-sauropod sauropodomorphs
Canardia
2013
Marnes d'Auzas Formation (Late Cretaceous , Maastrichtian )
France
May have been a close relative of Aralosaurus
Cardiodon
1841
Forest Marble Formation (Middle Jurassic , Bathonian )
England
The first sauropod ever named.[ 18] Known only from a tooth
Ceratosuchops
2021
Wessex Formation (Early Cretaceous , Barremian )
England
Differs from Baryonyx in subtle details of its skull
Cetiosauriscus
1927
Oxford Clay (Middle Jurassic , Callovian )
England
Has been assigned to a variety of positions around Eusauropoda[ 19] [ 20] [ 21]
Cetiosaurus
1841
Rutland Formation (Middle Jurassic , Bajocian to Bathonian )
England
The first sauropod known from decent remains. Once believed to be a large seagoing animal
Chondrosteosaurus
1876
Wessex Formation (Early Cretaceous , Barremian )
England
The air sacs in its vertebrae were originally believed to be filled with cartilage
Compsognathus
1859
Alcobaça Formation ?, Canjuers Lagerstätte , Solnhofen Limestone (Late Jurassic , Tithonian )
France Germany Portugal ?
One of the smallest known non-avian dinosaurs
Comptonatus
2024
Wessex Formation (Early Cretaceous , Barremian to Aptian )
England
One of the most completely known ornithopod taxa from the Isle of Wight
Concavenator
2010
La Huérguina Formation (Early Cretaceous , Barremian )
Spain
Preserves bumps on its ulna which have been interpreted as quill knobs,[ 22] although they might have been muscle attachments instead[ 23]
Craspedodon
1883
Glauconie de Lonzée Formation (Late Cretaceous , Santonian )
Belgium
May be the first neoceratopsian known from Europe[ 24]
Craterosaurus
1874
Woburn Sands Formation (Early Cretaceous , Aptian to Albian )
England
Potentially synonymous with Regnosaurus
Cruxicheiros
2010
Chipping Norton Limestone (Middle Jurassic , Bathonian )
England
Inconsistent in phylogenetic placement
Cryptosaurus
1869
Ampthill Clay (Late Jurassic , Oxfordian )
England
Only known from a single femur
Cumnoria
1888
Kimmeridge Clay (Late Jurassic , Kimmeridgian )
England
May be a species of Camptosaurus [ 25] or something a little more basal[ 26]
Dacentrurus
1902
Alcobaça Formation , Argiles d'Octeville , Kimmeridge Clay , Lourinhã Formation , Villar del Arzobispo Formation (Late Jurassic to Early Cretaceous , Kimmeridgian to Berriasian )
England France Portugal Spain
Known from abundant remains. Lived in a broad range
Demandasaurus
2011
Castrillo de la Reina Formation (Early Cretaceous , Barremian to Aptian )
Spain
Most closely related to African rebbachisaurids, suggesting a faunal exchange[ 27]
Dinodocus
1884
Lower Greensand Group (Early Cretaceous , Barremian )
England
The only known humerus is almost complete, missing only small portions
Dolichosuchus
1932
Löwenstein Formation (Late Triassic , Norian )
Germany
Originally classified as a stem-crocodile
Dornraptor
2024
Blue Lias ?/Charmouth Mudstone Formation ? (Early Jurassic , Sinemurian )
England
Originally thought to belong to Scelidosaurus ,[ 28] its remains were then informally named "Merosaurus" before their formal description in 2024[ 29]
Draconyx
2001
Lourinhã Formation (Late Jurassic , Tithonian )
Portugal
May have been a member of an ornithopod clade that did not appear in North America, unlike other groups of Late Jurassic animals[ 30]
Dracopelta
1980
Lourinhã Formation (Late Jurassic , Tithonian )
Portugal
The structure of its limbs suggests it might have had a cursorial lifestyle[ 31]
Dracoraptor
2016
Blue Lias (Early Jurassic , Hettangian )
Wales
May have been a shore dweller due to its island habitat[ 32]
Dromaeosauroides
2003
Jydegaard Formation (Early Cretaceous , Berriasian )
Denmark
A referred coprolite has been found which contains fish remains, but it could instead belong to a turtle[ 33]
Dubreuillosaurus
2005
Calcaire de Caen (Middle Jurassic , Bathonian )
France
Would have lived in a coastal mangrove swamp
Duriatitan
2010
Kimmeridge Clay (Late Jurassic , Kimmeridgian )
England
Originally named as a species of Cetiosaurus
Duriavenator
2008
Inferior Oolite (Middle Jurassic , Bajocian )
England
The basalmost known member of the Megalosaurinae, which aligns with its stratigraphic position[ 34]
Echinodon
1861
Purbeck Group (Early Cretaceous , Berriasian )
England
Originally misidentified as a herbivorous lizard
Efraasia
1973
Löwenstein Formation (Late Triassic , Norian )
Germany
Some remains assigned to this genus were originally classified in separate genera
Emausaurus
1990
Ciechocinek Formation (Early Jurassic , Toarcian )
Germany
One of the oldest and basalmost thyreophorans
Eotyrannus
2001
Wessex Formation (Early Cretaceous , Barremian )
England
Possessed grasping hands with three long fingers
Eousdryosaurus
2014
Lourinhã Formation (Late Jurassic , Kimmeridgian )
Portugal
Described as a dryosaurid[ 35] but one study suggests a close relationship with elasmarians[ 36]
Erectopus
1923
La Penthiève Beds (Early Cretaceous , Albian )
France
One of the youngest known European carnosaurs
Eucamerotus
1872
Wessex Formation (Early Cretaceous , Barremian )
England
Preserves extensive evidence of pneumatization
Eucercosaurus
1879
Cambridge Greensand (Early Cretaceous , Albian )
England
Sometimes considered an ankylosaur but one study assigns it to Iguanodontia[ 37]
Euronychodon
1991
Argiles et sables de Taveiro (Late Cretaceous , Campanian to Maastrichtian )
Portugal
Only known from teeth. Another species has been found in Uzbekistan
Europasaurus
2006
Süntel Formation (Late Jurassic , Kimmeridgian )
Germany
Much smaller than other sauropods due to its isolated island habitat [ 38]
Europatitan
2017
Castrillo de la Reina Formation (Early Cretaceous , Barremian to Aptian )
Spain
Some of this genus' remains include several vertebrae. The specific name, eastwoodi , honors director Clint Eastwood
Europelta
2013
Escucha Formation (Early Cretaceous , Albian )
Spain
Almost the entire skeleton is known
Eustreptospondylus
1964
Oxford Clay (Middle Jurassic to Late Jurassic , Callovian to Oxfordian )
England
May have swum between islands similar to a Komodo dragon [ 39]
Fylax
2021
Figuerola Formation (Late Cretaceous , Maastrichtian )
Spain
Lived very late despite its comparatively basal position[ 40]
Galvesaurus
2005
Villar del Arzobispo Formation (Late Jurassic , Tithonian )
Spain
Also spelled Galveosaurus . Two sets of paleontologists named the same fossil seemingly unaware of each other's work, although there is evidence that one of them had plaigiarized the others, but misspelled the name[ 41]
Garrigatitan
2021
Argiles et Grès à Reptiles Formation (Late Cretaceous , Campanian )
France
Known from remains of both adults and subadults
Garumbatitan
2023
Arcillas de Morella Formation (Early Cretaceous , Barremian )
Spain
Had a reduced claw on the third toe
Genusaurus
1995
Bevons Beds (Early Cretaceous , Albian )
France
Has been suggested to be either a noasaurid[ 42] or an abelisaurid[ 43]
Gideonmantellia
2012
Camarillas Formation (Early Cretaceous , Barremian )
Spain
Originally misidentified as a specimen of Hypsilophodon
Gigantosaurus
1869
Kimmeridge Clay (Late Jurassic , Kimmeridgian )
England
May have possessed osteoderms
Haestasaurus
2015
Wealden Group (Early Cretaceous , Berriasian to Valanginian )
England
Preserves impressions of differently-sized hexagonal scales
Halticosaurus
1908
Löwenstein Formation (Late Triassic , Norian )
Germany
Historically conflated with the bones of unrelated animals
Heptasteornis
1975
Sânpetru Formation (Late Cretaceous , Maastrichtian )
Romania
Once believed to be a giant prehistoric owl
Hesperonyx
2024
Lourinhã Formation (Late Jurassic , Kimmeridgian )
Portugal
Only known from the remains of the fore- and hindlimbs
Histriasaurus
1998
Unnamed formation (Early Cretaceous , Hauterivian to Barremian )
Croatia
Despite being discovered in Europe, it may have lived between southern Europe and Africa in life[ 44]
Horshamosaurus
2015
Weald Clay (Early Cretaceous , Barremian )
England
A supposed tibia has been reinterpreted as an ischium[ 45]
Hungarosaurus
2005
Csehbánya Formation (Late Cretaceous , Santonian )
Hungary
Possessed an elevated shoulder which may be an adaptation to high-browsing[ 46]
Hylaeosaurus
1833
Bückeberg Formation ?, Grinstead Clay Formation , Tunbridge Wells Sand Formation (Early Cretaceous , Valanginian )
England Germany ?
One of the three animals originally used to define the Dinosauria, along with Iguanodon and Megalosaurus
Hypselosaurus
1869
Argiles et Grès à Reptiles Formation (Late Cretaceous , Maastrichtian )
France
Several spherical eggs have been attributed to this taxon
Hypselospinus
2010
Wadhurst Clay Formation (Early Cretaceous , Valanginian )
England
Had elongated neural spines projecting from the top of its vertebrae
Hypsilophodon
1869
Wessex Formation (Early Cretaceous , Hauterivian to Barremian )
England
May have been a deer-like low browser that fed on young shoots and roots
Iberospinus
2022
Papo Seco Formation (Early Cretaceous , Barremian )
Portugal
Basal yet already displays some adaptations for a semiaquatic lifestyle
Iguanodon
1825
Arcillas de Morella Formation , Camarillas Formation , Sainte-Barbe Clays Formation , Wadhurst Clay Formation , Weald Clay , Wealden Group , Wessex Formation (Early Cretaceous , Barremian to Aptian )
Belgium England Germany ? Spain
Multiple remains are known which make it one of the best known dinosaurs
Iliosuchus
1932
Taynton Limestone Formation (Middle Jurassic , Bathonian )
England
Only known from three ilia
Iuticosaurus
1993
Upper Greensand Formation , Wessex Formation (Early Cretaceous , Hauterivian to Barremian )
England
Two species have been named, each from a single caudal vertebra
Juratyrant
2013
Kimmeridge Clay (Late Jurassic , Tithonian )
England
Originally named as a species of Stokesosaurus
Juravenator
2006
Painten Formation (Late Jurassic , Kimmeridgian )
Germany
Impressions of both scales and feathers are known. The tail preserves structures that may be integumentary sense organs like those of crocodiles, which it may have used to prey on fish at night[ 47]
Lexovisaurus
1957
Oxford Clay (Middle Jurassic , Callovian )
England
Its pelvis was greatly enlarged
Liliensternus
1984
Trossingen Formation (Late Triassic , Norian to Rhaetian )
Germany
Although commonly depicted with a head crest, there is no evidence for such a feature
Lirainosaurus
1999
Marnes Rouges Inférieures Formation ?, Sierra Perenchiza Formation , Sobrepena Formation (Late Cretaceous , Campanian to Maastrichtian )
France ? Spain
For a titanosaur, it was small and had a relatively gracile build
Lohuecotitan
2016
Villalba de la Sierra Formation (Late Cretaceous , Campanian to Maastrichtian )
Spain
May have had "bulb-and-root"-type osteoderms, which are abundant at the type locality[ 48]
Lophostropheus
2007
Moon-Airel Formation (Late Triassic to Early Jurassic , Rhaetian to Hettangian )
France
The only substantially well-known theropod from the Triassic-Jurassic boundary[ 49]
Loricatosaurus
2008
Marnes à Belemnopsis latesulcatus Formation , Oxford Clay (Middle Jurassic , Callovian )
England France
Had narrow, flat plates on its back and round, pointed spines that ran along the tail
Losillasaurus
2001
Villar del Arzobispo Formation (Late Jurassic to Early Cretaceous ?, Kimmeridgian to Berriasian ?)
Spain
Was heterodont , having four types of teeth, one of which was heart-shaped[ 50]
Lourinhanosaurus
1998
Lourinhã Formation (Late Jurassic , Kimmeridgian to Tithonian )
Portugal
More than one hundred eggs have been referred to this taxon
Lourinhasaurus
1998
Lourinhã Formation (Late Jurassic , Kimmeridgian )
Portugal
Closely related to Camarasaurus but with proportionately longer forelimbs
Lusotitan
2003
Lourinhã Formation (Late Jurassic , Tithonian )
Portugal
Originally named as a European species of Brachiosaurus
Lusovenator
2020
Lourinhã Formation (Late Jurassic to Early Cretaceous , Kimmeridgian to Berriasian )
Portugal
The oldest carcharodontosaurian known from Eurasia
Macrurosaurus
1869
Cambridge Greensand , Chalk Group ? (Early Cretaceous , Albian )
England
Only known from a series of caudal vertebrae
Magnamanus
2016
Golmayo Formation (Early Cretaceous , Hauterivian to Barremian )
Spain
Possessed relatively enlarged hands
Magnosaurus
1932
Inferior Oolite (Middle Jurassic , Bajocian )
England
Confusingly, a referred specimen was simultaneously named as a species of this genus and of Sarcosaurus
Magyarosaurus
1932
Sânpetru Formation (Late Cretaceous , Maastrichtian )
Romania
An insular dwarf titanosaur that was one of the smallest of its group
Mantellisaurus
2007
Arcillas de Morella Formation , Lower Greensand Group , Sainte-Barbe Clays Formation , Vectis Formation , Wessex Formation (Early Cretaceous , Barremian to Aptian )
Belgium England Germany ? Spain
Several specimens are known. Distinguishable from the contemporary Iguanodon by its more gracile build
Marmarospondylus
1875
Forest Marble Formation (Middle Jurassic , Bathonian )
England
Usually assigned to the genus Bothriospondylus , but this cannot be confirmed
Matheronodon
2017
Argiles et Grès à Reptiles Formation (Late Cretaceous , Campanian )
France
Had extremely specialized dentition that may have been an adaptation to feeding on tough monocot plants[ 51]
Megalosaurus
1824
Chipping Norton Limestone , Taynton Limestone Formation (Middle Jurassic , Bathonian )
England
The first non-avian dinosaur scientifically named and described
Metriacanthosaurus
1964
Oxford Clay (Late Jurassic , Oxfordian )
England
Possessed relatively tall neural spines for a carnosaur
Miragaia
2009
Lourinhã Formation (Late Jurassic , Kimmeridgian to Tithonian )
Portugal
Had an extremely elongated neck made up of seventeen vertebrae. May be synonymous with Dacentrurus [ 52] [ 53]
Mochlodon
1881
Csehbánya Formation , Gosau Group (Late Cretaceous , Santonian to Campanian )
Austria Hungary
The type specimen was originally named as a species of Iguanodon
Morelladon
2015
Arcillas de Morella Formation (Early Cretaceous , Barremian )
Spain
Possessed a low sail on its back supported by elongated neural spines
Morinosaurus
1874
Unnamed formation (Late Jurassic , Kimmeridgian )
France
Poorly known
Neosodon
1885
Sables et Grès à Trigonia gibbosa (Late Jurassic , Tithonian )
France
No species are assigned to this genus. Popularly associated with "Iguanodon " praecursor but is in fact a separate taxon[ 54]
Neovenator
1996
Wessex Formation (Early Cretaceous , Hauterivian to Barremian )
England
Had a complex series of neurovascular canals lining its snout[ 55]
Normanniasaurus
2013
Poudingue Ferrugineux (Early Cretaceous , Albian )
France
Represents a European radiation of basal titanosaurs[ 56]
Notatesseraeraptor
2019
Klettgau Formation (Late Triassic , Norian )
Switzerland
Combines features of different groups of basal theropods
Nuthetes
1854
Lulworth Formation (Early Cretaceous , Berriasian )
England
Only known from jaws and teeth. Traditionally identified as a dromaeosaurid but it might be a proceratosaurid instead[ 57]
Oblitosaurus
2023
Villar del Arzobispo Formation (Late Jurassic , Kimmeridgian to Tithonian )
Spain
The largest ornithopod known from the Late Jurassic of Europe
Oceanotitan
2019
Lourinhã Formation (Late Jurassic , Kimmeridgian )
Portugal
Potentially the oldest known somphospondylian[ 58]
Ohmdenosaurus
1978
Posidonia Shale (Early Jurassic , Toarcian )
Germany
Originally misidentified as a plesiosaur
Oplosaurus
1852
Wessex Formation (Early Cretaceous , Barremian )
England
The holotype tooth was pointed, which led to its misidentification as a carnivorous reptile[ 59]
Ornithodesmus
1887
Wessex Formation (Early Cretaceous , Barremian )
England
Historically conflated with the remains of the pterosaur Istiodactylus
Ornithopsis
1870
Wealden Group (Early Cretaceous , Barremian )
England )
Originally believed to be an intermediate form between birds, pterosaurs and dinosaurs
Orthomerus
1883
Maastricht Formation (Late Cretaceous , Maastrichtian )
Netherlands
Potentially dubious and undiagnostic[ 60]
Ostromia
2017
Painten Formation (Late Jurassic , Tithonian )
Germany
Considered a small pterosaur until it was redescribed as a specimen of Archaeopteryx in 1970.[ 61] Some of its features are similar to those of Anchiornis
Owenodon
2009
Lulworth Formation (Early Cretaceous , Berriasian )
England
Has been assigned to Iguanodon and Camptosaurus before it received its own genus
Paludititan
2010
Sânpetru Formation (Late Cretaceous , Maastrichtian )
Romania
Some of its bones are identical to those of Magyarosaurus , but their synonymy cannot be confirmed
Pararhabdodon
1993
Tremp Formation (Late Cretaceous , Maastrichtian )
Spain
The first lambeosaurine identified from Europe
Pareisactus
2019
Tremp Formation (Late Cretaceous , Maastrichtian )
Spain
Represented by a single scapula recovered from a hadrosaur bonebed
Pelecanimimus
1994
La Huérguina Formation (Early Cretaceous , Barremian )
Spain
Preserves extensive soft tissue impressions revealing the presence of a keratinous head crest and a pelican-like gular pouch
Pelorosaurus
1850
Tunbridge Wells Sand Formation (Early Cretaceous , Hauterivian )
England
The first sauropod identified as a terrestrial animal
Pendraig
2021
Pant-y-Ffynnon Quarry (Late Triassic , Rhaetian )
Wales
Would have lived on a dry limestone island[ 62]
Phyllodon
1973
Alcobaça Formation (Late Jurassic , Kimmeridgian )
Portugal
The front and back of its teeth were asymmetrical
Piveteausaurus
1977
Marnes de Dives (Middle Jurassic , Callovian )
France
Has been suggested to be a species of Proceratosaurus
Plateosaurus
1837
Klettgau Formation , Löwenstein Formation , Lunde Formation , Trossingen Formation (Late Triassic , Norian )
France ? Germany Norway Switzerland
Known from over a hundred skeletons, several of them nearly complete
Pneumatoraptor
2010
Csehbánya Formation (Late Cretaceous , Santonian )
Hungary
One study suggests a position as a possible early palaeognath[ 63]
Poekilopleuron
1836
Calcaire de Caen (Middle Jurassic , Bathonian )
France
Its holotype was found alongside fish remains[ 64]
Polacanthus
1865
Wessex Formation (Early Cretaceous , Barremian )
England
Possessed a flat sacral shield dotted with small bumps
Portellsaurus
2021
Margas de Mirambell Formation (Early Cretaceous , Barremian )
Spain
Closely related to Ouranosaurus [ 65]
Priodontognathus
1875
Lower Calcareous Grit (Late Jurassic , Oxfordian )
England
Only known from a single maxilla with teeth
Proa
2012
Escucha Formation (Early Cretaceous , Albian )
Spain
The tip of its jaw was shaped like the bow of a ship
Proceratosaurus
1926
Great Oolite Group (Middle Jurassic , Bathonian )
England
Preserves a small horn on its snout which may have anchored a crest as in the related Guanlong
Procompsognathus
1913
Löwenstein Formation (Late Triassic , Norian )
Germany
Has been suggested to be non-dinosaurian
Protathlitis
2023
Arcillas de Morella Formation (Early Cretaceous , Barremian )
Spain
Large but basal for a spinosaurid
Pterospondylus
1913
Trossingen Formation (Late Triassic , Norian )
Germany
Known from only a single, large vertebra
Pyroraptor
2000
Argiles et Grès à Reptiles Formation (Late Cretaceous , Campanian to Maastrichtian )
France
Its holotype specimen was exposed by a forest fire
Qunkasaura
2024
Villalba de la Sierra Formation (Late Cretaceous , Campanian to Maastrichtian )
Spain
May have been closely related to Abditosaurus [ 66]
Regnosaurus
1848
Tunbridge Wells Sand Formation (Early Cretaceous , Berriasian to Valanginian )
England
Potentially a stegosaur similar to Huayangosaurus [ 67]
Rhabdodon
1869
Grès de Labarre , Gres de Saint-Chinian , Marnes Rouges Inférieures Formation , Villalba de la Sierra Formation (Late Cretaceous , Maastrichtian )
France Spain
Although most rhabdodontids are insular dwarfs, this genus may represent an instance of island gigantism as it is much larger than other members of its family[ 68]
Riabininohadros
2020
Unnamed formation (Late Cretaceous , Maastrichtian )
Crimea ( Ukraine de jure )
Possessed a femur so unique it has no morphological equivalents across all of Iguanodontia
Riojavenatrix
2024
Enciso Group (Early Cretaceous , Barremian to Aptian )
Spain
Originally identified as a specimen of Baryonyx
Riparovenator
2021
Wessex Formation (Early Cretaceous , Barremian )
England
Had elongated spines projecting from its caudal vertebrae somewhat similar to those of Spinosaurus
Ruehleia
2001
Trossingen Formation (Late Triassic , Norian )
Germany
Known from a single, nearly complete skeleton
Saltriovenator
2018
Saltrio Formation (Early Jurassic , Sinemurian )
Italy
The biggest theropod from the Early Jurassic yet known
Sarcolestes
1893
Oxford Clay (Middle Jurassic , Callovian )
England
Originally misidentified as a carnivorous dinosaur
Sarcosaurus
1921
Scunthorpe Mudstone (Early Jurassic , Hettangian to Sinemurian )
England
Has been variously suggested to be in different positions at the base of Neotheropoda[ 69] [ 70] [ 71] [ 72]
Scelidosaurus
1859
Charmouth Mudstone Formation (Early Jurassic , Sinemurian to Pliensbachian )
England
Carried hundreds of small osteoderms in several rows along its back
Schleitheimia
2020
Klettgau Formation (Late Triassic , Norian )
Switzerland
Possessed a relatively enlarged ilium
Scipionyx
1998
Pietraroja Plattenkalk (Early Cretaceous , Albian )
Italy
So well preserved that several internal organs and their positions in life could be accurately reconstructed
Sciurumimus
2012
Torleite Formation (Late Jurassic , Kimmeridgian )
Germany
Had a bushy tail similar to that of a squirrel
Soriatitan
2017
Golmayo Formation (Early Cretaceous , Hauterivian to Barremian )
Spain
The first confirmed brachiosaurid known from Early Cretaceous Europe
Stenopelix
1857
Obernkirchen Sandstein (Early Cretaceous , Berriasian )
Germany
Possibly closely related to basal ceratopsians from Late Jurassic China[ 73]
Streptospondylus
1832
Marnes de Dives ?/Marnes de Villers ? (Middle Jurassic to Late Jurassic , Callovian to Oxfordian )
France
Originally believed to represent a marine crocodile
Struthiosaurus
1871
Argiles et Grès à Reptiles Formation , Gosau Group , Sânpetru Formation (Late Cretaceous , Santonian to Maastrichtian )
Austria France Hungary ? Romania
Analysis of its braincase suggests poor hearing and a sluggish, solitary lifestyle[ 74]
Syngonosaurus
1879
Cambridge Greensand (Early Cretaceous , Albian )
England
Usually considered a synonym of Acanthopholis but it has been reinterpreted as an iguanodont[ 37]
Tamarro
2021
Tremp Formation (Late Cretaceous , Maastrichtian )
Spain
May have been closely related to Asian troodontids[ 75]
Tanystrosuchus
1963
Löwenstein Formation (Late Triassic , Norian )
Germany
The only known vertebra was once misidentified as a phytosaur
Tarascosaurus
1991
Fuvelian Beds (Late Cretaceous , Campanian )
France
If an abelisaurid it would be one of the few northern members of the group
Tastavinsaurus
2008
Forcall Formation , Xert Formation (Early Cretaceous , Barremian )
Spain
Could represent an obscure group of macronarians called Laurasiformes[ 76]
Teinurosaurus
1928
Mont-Lambert Formation (Late Jurassic , Tithonian )
France
Poorly known
Telmatosaurus
1903
Sânpetru Formation (Late Cretaceous , Maastrichtian )
Romania
One specimen preserves a facial deformity caused by an ameloblastoma [ 77]
Tethyshadros
2009
Calcare di Aurisina (Late Cretaceous , Campanian )
Italy
Had limbs adapted for high speed but were too short for running[ 46]
Thecocoelurus
1923
Wessex Formation (Early Cretaceous , Barremian )
England
Has been assigned to a variety of theropod groups throughout history
Thecodontosaurus
1836
Magnesian Conglomerate (Late Triassic , Rhaetian )
England
Could have been an extremely fast bipedal runner[ 78]
Thecospondylus
1882
Wealden Group (Early Cretaceous , Valanginian to Hauterivian )
England
Indeterminate within Dinosauria
Torvosaurus
1979
Kimmeridge Clay ?, Lourinhã Formation , Ornatenton Formation , Vega Formation ? (Middle Jurassic to Late Jurassic , Callovian to Tithonian )
England ? Germany Portugal Spain ?
The type species was found in the United States. Several species lived around the world, including in Europe, but T. gurneyi is the only non-North American species to receive a formal name. It represents Europe's largest terrestrial predator[ 79]
Transylvanosaurus
2022
Pui Beds (Late Cretaceous , Maastrichtian )
Romania
Had an unusually wide skull compared to other rhabdodontids
Trimucrodon
1973
Lourinhã Formation (Late Jurassic , Kimmeridgian )
Portugal
Possessed wedge-shaped teeth
Tuebingosaurus
2022
Trossingen Formation (Late Triassic , Norian )
Germany
Although originally assigned to Plateosaurus , it contains several features that point to a more derived position
Turiasaurus
2006
Villar del Arzobispo Formation (Late Jurassic , Kimmeridgian to Tithonian )
Spain
Extremely large despite not being a member of Neosauropoda
Valdosaurus
1977
Weald Clay , Wealden Group , Wessex Formation (Early Cretaceous , Berriasian to Barremian )
England
Large and similar to Dryosaurus
Vallibonavenatrix
2019
Arcillas de Morella Formation (Early Cretaceous , Barremian )
Spain
One of the most complete spinosaurids known from Iberia
Variraptor
1998
Argiles et Grès à Reptiles Formation (Late Cretaceous , Campanian to Maastrichtian )
France
May be closely related to Bambiraptor [ 63]
Vectaerovenator
2020
Lower Greensand Group (Early Cretaceous , Aptian )
England
Only known from four vertebrae but are distinct enough to be classified as their own genus
Vectidromeus
2023
Wessex Formation (Early Cretaceous , Barremian )
England
Originally described as a relative of the coeval Hypsilophodon , but a subsequent analysis suggests possible dryosaurid affinities for this genus[ 80]
Vectipelta
2023
Wessex Formation (Early Cretaceous , Hauterivian to Barremian )
England
May have been more closely related to Asian ankylosaurs than to European ones[ 81]
Vectiraptor
2021
Wessex Formation (Early Cretaceous , Barremian )
England
Shares some features with North American dromaeosaurids
Velocipes
1932
Lissauer Breccia (Late Triassic , Norian )
Poland
Has been considered a dubious, indeterminate vertebrate[ 82]
Volgatitan
2018
Unnamed formation (Early Cretaceous , Hauterivian )
Russia
Closely related to South American titanosaurs
Vouivria
2017
Calcaires de Clerval (Late Jurassic , Oxfordian )
France
The oldest known titanosauriform
Wiehenvenator
2016
Ornatenton Formation (Middle Jurassic , Callovian )
Germany
Before its formal description, it had been nicknamed the "Minden Monster"
Xenoposeidon
2007
Wealden Group (Early Cretaceous , Berriasian to Valanginian )
England
Only known from a single, very unique vertebra
Yaverlandia
1971
Vectis Formation ?, Wessex Formation (Early Cretaceous , Barremian )
England
Originally misidentified as a pachycephalosaur because of its thick skull roof
Zalmoxes
2003
Densuş-Ciula Formation , Sânpetru Formation , Sebeş Formation (Late Cretaceous , Maastrichtian )
Romania
Two species of contrasting sizes have been named
Zby
2014
Lourinhã Formation (Late Jurassic , Kimmeridgian )
Portugal
Originally believed to be a specimen of Turiasaurus
Invalid and potentially valid genera
Agrosaurus macgillivrayi : Originally mistakenly thought to be from Australia. It is now thought to be more likely from England and possibly a synonym of Thecodontosaurus .
Archaeopteryx : A well-known taxon that combines bird-like pennaceous feathers with the teeth, claws and long tail of reptiles. It is usually considered a basal avialan but it might also be a non-avian deinonychosaur closely related to dromaeosaurids.
Balaur bondoc : A strange paravian that possessed a suite of unique features, such as robust muscles, two sickle claws, a didactyl manus and a deep gut. It may have been an island-dwelling herbivore or omnivore. Studies show it either as a velociraptorine dromaeosaurid or a basal avialan. It may be a synonym of Elopteryx .
"Bihariosaurus bauxiticus ": Although sometimes presented as a valid taxon, it is actually a nomen nudum .
Ceratosaurus : Some specimens, mostly teeth, from Portugal and Switzerland have been assigned to this genus, but not to a specific species.
Darwinsaurus evolutionis : May be a synonym of either Hypselospinus or Mantellisaurus .
Delapparentia turolensis : Said to be indistinguishable from Iguanodon .
Dinheirosaurus lourinhanensis : Possibly a second species of Supersaurus .
Elopteryx nopcsai : Has been placed in different maniraptoran groups throughout history. A recent study suggests an avialan position for the taxon based on new fossil material.
Gresslyosaurus : Often thought to be synonymous with Plateosaurus , although several differences between them have been noted.
Huxleysaurus hollingtoniensis : Potentially a synonym of Hypselospinus .
"Ischyrosaurus ": The generic name Ischyrosaurus is preoccupied. The dinosaur may be a synonym of Ornithopsis .
Koutalisaurus kohlerorum : Usually seen as a synonym of Pararhabdodon , but it could also be its own taxon.
Kukufeldia tilgatensis : May be a synonym of Barilium .
"Liassaurus huenei ": Could potentially be a second specimen of Sarcosaurus .
Lusitanosaurus liasicus : Although originally described as an early thyreophoran, later research has questioned this assignment and considered it a non-dinosaurian archosauromorph.
Mantellodon carpenteri : Known from a single specimen from Maidstone affectionally nicknamed a "Mantell -piece". While originally thought to be a specimen of Iguanodon , it may in fact be a synonym of Mantellisaurus .
"Newtonsaurus " cambrensis : Said to be an early theropod, but it may be indeterminate within Archosauria.
Oligosaurus adelus : May be synonymous with Mochlodon .
Ornithomerus gracilis : May be synonymous with Mochlodon .
Pachysauriscus ajax : Usually seen as a synonym of Plateosaurus , but a 2023 review considered it distinct.
Pantydraco caducus : Originally named as a species of Thecodontosaurus and indeed it may belong to that genus.
Polacanthoides ponderosus : Potentially a synonym of Hylaeosaurus , Polacanthus or both (a chimera).
Proplanicoxa galtoni : May be a junior synonym of Mantellisaurus .
Rhadinosaurus alcimus : Suggested to be synonymous with Struthiosaurus , but it might have been a crocodilian instead.
Sellacoxa pauli : Possibly synonymous with Barilium .
Stegosaurus : Some bones from Portugal have been assigned to this genus, but not to a particular species.
Suchosaurus : Two species have been named, both from teeth. They are indistinguishable from those of baryonychines and may in fact belong to Baryonyx .
Taveirosaurus costai : Only known from teeth. Initially thought to be an ornithischian but it might be a eutriconodont mammal instead.
Therosaurus : This genus was named to accommodate the original type species of Iguanodon , I. anglicus . However, it was named before the type species was transferred to I. bernissartensis , so it remains a junior synonym of Iguanodon .
Valdoraptor oweni : Potentially a junior synonym of Thecocoelurus .
Wellnhoferia grandis : May be another specimen of Archaeopteryx .
Timeline
This is a timeline of selected dinosaurs from the list above. Time is measured in Ma , megaannum , along the x-axis.
See also
Notes
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