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Canis nehringi; Temporal range: Late Pleistocene (Lujanian); ~0.8–0.011 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Carnivora
Family:	Canidae
Genus:	Canis
Species:	†C. nehringi
Binomial name
	†Canis nehringi; Ameghino 1902

Canis nehringi is an extinct species of canid. Canis gezi, a poorly known small wolf from the Ensenadan of South America, appears to have given rise to Canis nehringi, a Lujanian species from Argentina. The species was first described by Florentino Ameghino in 1902.^[1]

Canis gezi

Canis dirus made its appearance in South America in the late Pleistocene, and seems to have been restricted to the north and west coasts. Its remains have not been found in the area of Argentina that produced Canis gezi and Canis nehringi, and their remains have not been found elsewhere in South America.^[3]^: 174–5 Some researchers have proposed that Canis dirus may have originated in South America.^[4]^: 116^[5]^[6] In 1988, a study of these two large South American wolves described them with Canis gezi found in South American Ensenadan deposits that relate to the North American late Blancan and Irvingtonian, and Canis nehringi found in South American Lujanian deposits of the Lujan Formation that relate to the Late Pleistocene. Given their similarities and timeframes, it was proposed that Canis gezi was the ancestor of Canis nehringi.^[7]^: 113

The study indicated that Canis gezi was most similar to the late Irvingtonian Aenocyon dirus nebrascensis (proposed early Dire wolf) and was its sister taxa,^[7]^: 55 but Canis nehringi had a closer relationship to Rancholabrean Canis dirus.^[7]^: 113 The study found that Canis dirus was the most derived genus Canis species in the New World, and compared to Canis nehringi was larger in size and construction of its lower molars that were increased for more efficient predation.^[7]^: 113 In 2009, Tedford proposed that because there was now seen a link between Canis armbrusteri and the Rancholabrean Canis dirus, that a case could be argued for a collateral South American lineage linking Canis gezi with Canis nehringi. These two clades share dental and cranial similarities developed for hypercarnivory, suggesting a common ancestor for both clades.^[3]^: 148

In 2010, a study found that DNA analysis and the dental characteristics of South America hypercanivorous canids showed a "South American clade" and the Canis clade. Canis gezi was a member of the "South American clade" of carnivores, but Canis dirus and Canis nehringi were included as derived species in the Canis clade. Canis dirus was the sister taxon of Canis lupus, but the scientific scoring used in this study for Canis nehringi was identical to those observed in Canis dirus, which supports the proposal that both could have been the same species.^[8]^: 472

In 2018, a study found that Canis gezi did not fall under genus Canis and should be classified under the subtribe Cerdocyonina, however no genus was proposed.^[9]

References

^ Prevosti et al., 2004, p.32
^ Artwork by Sergio De la Rosa Martinez (refer to the Summary of this pix)
^ ^a ^b Tedford, Richard H.; Wang, Xiaoming; Taylor, Beryl E. (2009). "Phylogenetic Systematics of the North American Fossil Caninae (Carnivora: Canidae)" (PDF). Bulletin of the American Museum of Natural History. 325: 1–218. doi:10.1206/574.1. S2CID 83594819.
^ R. M. Nowak. 1979. North American Quaternary Canis. Monograph of the Museum of Natural History, University of Kansas 6:1-154, page 106 LINK:[1]
^ Dundas, R.G. (1999). "Quaternary records of the dire wolf, Canis dirus, in North and South America" (PDF). Boreas. 28 (3): 375–385. Bibcode:1999Borea..28..375D. doi:10.1111/j.1502-3885.1999.tb00227.x. S2CID 129900134. Archived from the original (PDF) on 2020-09-27. Retrieved 2016-02-12.
^ B. Kurten and E. Anderson. 1980. Pleistocene mammals of North America 1-442
^ ^a ^b ^c ^d Berta, A. 1988. Quaternary evolution and biogeography of the large South American Canidae (Mammalia: Carnivora). University of California Publications in Geological Sciences 132:1–49.
^ Prevosti, Francisco J. (2010). "Phylogeny of the large extinct South American Canids (Mammalia, Carnivora, Canidae) using a "total evidence" approach". Cladistics. 26 (5): 456–481. doi:10.1111/j.1096-0031.2009.00298.x. PMID 34875763. S2CID 86650539.
^ Zrzavý, Jan; Duda, Pavel; Robovský, Jan; Okřinová, Isabela; Pavelková Řičánková, Věra (2018). "Phylogeny of the Caninae (Carnivora): Combining morphology, behaviour, genes and fossils". Zoologica Scripta. 47 (4): 373–389. doi:10.1111/zsc.12293. S2CID 90592618.