Carolyn Widney "Carol" Greider (born April 15, 1961) is an American molecular biologist . She is Professor and Director of Molecular Biology and Genetics at Johns Hopkins University .[ 1]
Greider discovered the enzyme telomerase in 1984, when she was a graduate student of Elizabeth Blackburn at the University of California, Berkeley . Greider pioneered research on the structure of telomeres , the ends of the chromosomes . She was awarded the 2009 Nobel Prize for Physiology or Medicine, with Blackburn and Jack Szostak , for their discovery that telomeres are protected from progressive shortening by the enzyme telomerase.[ 2]
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