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Symphyotrichum spathulatum
Conservation status
	; Secure (NatureServe)
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Clade:	Angiosperms
Clade:	Eudicots
Clade:	Asterids
Order:	Asterales
Family:	Asteraceae
Tribe:	Astereae
Subtribe:	Symphyotrichinae
Genus:	Symphyotrichum
Subgenus:	Symphyotrichum subg. Symphyotrichum
Section:	Symphyotrichum sect. Occidentales
Species:	S. spathulatum
Binomial name
	Symphyotrichum spathulatum; (Lindl.) G.L.Nesom
Varieties
	List S. spathulatum var. spathulatum ; S. spathulatum var. intermedium (A.Gray) G.L.Nesom ; S. spathulatum var. yosemitanum (A.Gray) G.L.Nesom ;
	Native distribution
Synonyms
	Basionym; Aster spathulatus Lindl.; Species Aster aestivus Rothr. ; Aster ascendens var. delectabilis (H.M.Hall) Jeps. ; Aster ascendens var. fremontii Torr. & A.Gray ; Aster delectabilis H.M.Hall ; Aster fremontii A.Gray ; Aster fremontii f. pallidus Cockerell ; Aster fremontii var. parishii A.Gray ; Aster fremontii f. viridus Cockerell ; Aster occidentalis Torr. & A.Gray ; Aster occidentalis var. delectabilis (H.M.Hall) Ferris ; Aster occidentalis var. fremontii (Torr. & A.Gray) A.G.Jones ; Aster occidentalis var. parishii (A.Gray) Ferris ; Aster occidentalis var. scabriusculus A.Gray ; Aster occidentalis var. typicus Cronquist ; Symphyotrichum spathulatum var. fremontii (Torr. & A.Gray) G.L.Nesom ; Tripolium occidentale Nutt. ; ; ; Varieties var. intermedium Aster occidentalis var. intermedius A.Gray ; Aster occidentalis subsp. intermedius (A.Gray) Piper ; Aster spathulatus var. intermedius (A.Gray) Cronquist ; Aster vallicola Greene ; ; ; ; var. yosemitanum Aster ascendens var. yosemitanus A.Gray ; Aster occidentalis var. yosemitanus (A.Gray) Cronquist ; Aster paludicola Piper ; Aster yosemitanus Greene ; ; ; ;

Symphyotrichum spathulatum (formerly Aster spathulatus) is a species of flowering plant in the family Asteraceae native to western North America including northwestern Mexico. Commonly known as western mountain aster, it is a perennial, herbaceous plant that may reach 20 to 80 centimeters (8 to 31 inches) tall. Its flowers, which open in July and August, have violet ray florets and yellow disk florets.^[5]

Description

Symphyotrichum spathulatum blooms in July and August and is a colony-forming perennial that grows typically 1–5 hairless or mostly hairless stems from a long rhizome. It ranges from 20 to 80 centimeters (8 to 31 inches) in height and has thin, entire leaves with little to no hair that are 5 to 15 centimeters (2 to 6 inches) long. The leaves are linear or elliptical, narrow, and sometimes obovate at the base of the plant. The upper leaves are shorter at 3 to 6 centimeters (1+1⁄8 to 2+3⁄8 inches).^[5]

The flower heads grow in corymbiform to paniculiform arrays with little branching. The involucres are 5–10 millimeters (1⁄5–2⁄5 inch) and bell-shaped, and their phyllaries are in 3–5 series. There are 15–40 violet ray florets that are 9–15 millimeters (7⁄20–3⁄5 inch) long and 1–2 millimeters (1⁄20–1⁄10 inch) wide. These surround the flower centers composed of 30–80 (sometimes up to 100) yellow disk florets.^[5]

The seeds are brown, hairy cypselae 2.5–3.5 mm (1⁄10–3⁄20 in) long with about 4 nerves and white pappi that are 5–7 millimeters (1⁄5–3⁄10 inch) long.^[5]

Chromosomes

Symphyotrichum spathulatum has a base number of x = 8.^[6] Diploid, tetraploid, hexaploid, and octaploid cytotypes with respective chromosome counts of 16, 32, 48, and 64 have been reported, depending upon the infraspecies, as follows:

S. spathulatum var. spathulatum: 2n = 2x = 16, 2n = 4x = 32, 2n = 6x = 48, and 2n = 8x = 64^[7]
S. spathulatum var. intermedium: 2n = 4x = 32, 2n = 6x = 48, and 2n = 8x = 64^[8]
S. spathulatum var. yosemitanum: 2n = 2x = 16, 2n = 4x = 32^[9]

Taxonomy

Symphyotrichum spathulatum is one of the species within Symphyotrichum sect. Occidentales.^[a]^[11] S. spathulatum was first formally described by John Lindley in 1834 as Aster spathulatus.^[12]

Three varieties of Symphyotrichum spathulatum are recognized, including the autonym:^[5]

S. spathulatum var. spathulatum
S. spathulatum var. intermedium
S. spathulatum var. yosemitanum

S. spathulatum is one of the parents of the two allopolyploidal Symphyotrichum species S. ascendens^[13] and S. defoliatum.^[14]

Distribution and habitat

Symphyotrichum spathulatum is found in the western United States, western Canada, and northwestern Mexico. It is native to British Columbia and Alberta in Canada, and in the United States from Washington state east to Montana, south to New Mexico, west to California, and north again to Oregon. There is no recorded presence in Arizona. In Mexico, it is native to the states bordering the Gulf of California.^[2] It grows at 1,200–2,900 meters (3,900–9,500 feet) (or lower) in meadows on mountain slopes and open aspen and coniferous forests.^[7]

S. spathulatum var. intermedium is native to British Columbia, California, Idaho, Nevada, Oregon, and Washington state,^[3] only occasionally being found in California and Nevada. It grows at 1,200–2,200 m (3,900–7,200 ft) in grasslands and meadows on mountain slopes and in open coniferous forests.^[8]

S. spathulatum var. yosemitanum is restricted from southern Oregon to the Sierra Nevada of California. It can be found at 1,200–2,200 m (3,900–7,200 ft) in oak woodlands and coniferous forests.^[15]

Conservation

As of July 2021^[update], NatureServe listed Symphyotrichum spathulatum as Secure (G5) worldwide and Possibly Extirpated (SX) in Alberta.^[1] S. spathulatum var. intermedium and S. spathulatum var. spathulatum were reported as Secure Varieties (T5),^[16]^[17] and no status rank was given for S. spathulatum var. yosemitanum.^[18]

Notes

^ Occidentales is listed as a section in Flora of Missouri,^[10] whereas on Canadian botanist John C. Semple's Astereae Lab website, it is given as a subsection of section Symphyotrichum.^[11]

Citations

^ ^a ^b NatureServe 2021a.
^ ^a ^b ^c ^d ^e POWO 2021a.
^ ^a ^b POWO 2021b.
^ POWO 2021c.
^ ^a ^b ^c ^d ^e Brouillet et al. 2006a.
^ Semple n.d.a.
^ ^a ^b Brouillet et al. 2006c.
^ ^a ^b Brouillet et al. 2006b.
^ Allen 2012.
^ Yatskievych n.d.
^ ^a ^b Semple 2021a.
^ IPNI 2021.
^ Semple 2021b.
^ Semple n.d.b.
^ Brouillet et al. 2006d.
^ NatureServe 2021b.
^ NatureServe 2021c.
^ NatureServe 2021d.

References

Allen, G.A. (2012). "Symphyotrichum spathulatum var. yosemitanum". In Jepson Flora Project (ed.). Jepson eFlora. The Jepson Herbarium, University of California, Berkeley. Retrieved 23 September 2022.
Brouillet, L.; Semple, J.C.; Allen, G.A.; Chambers, K.L.; Sundberg, S.D. (2006a). "Symphyotrichum spathulatum". In Flora of North America Editorial Committee (ed.). Flora of North America North of Mexico (FNA). Vol. 20. New York and Oxford: Oxford University Press. Retrieved 23 July 2021 – via eFloras.org, Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
Brouillet, L.; Semple, J.C.; Allen, G.A.; Chambers, K.L.; Sundberg, S.D. (2006b). "Symphyotrichum spathulatum var. intermedium". In Flora of North America Editorial Committee (ed.). Flora of North America North of Mexico (FNA). Vol. 20. New York and Oxford: Oxford University Press. Retrieved 23 July 2021 – via eFloras.org, Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
Brouillet, L.; Semple, J.C.; Allen, G.A.; Chambers, K.L.; Sundberg, S.D. (2006c). "Symphyotrichum spathulatum var. spathulatum". In Flora of North America Editorial Committee (ed.). Flora of North America North of Mexico (FNA). Vol. 20. New York and Oxford: Oxford University Press. Retrieved 23 July 2021 – via eFloras.org, Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
Brouillet, L.; Semple, J.C.; Allen, G.A.; Chambers, K.L.; Sundberg, S.D. (2006d). "Symphyotrichum spathulatum var. yosemitanum". In Flora of North America Editorial Committee (ed.). Flora of North America North of Mexico (FNA). Vol. 20. New York and Oxford: Oxford University Press. Retrieved 23 July 2021 – via eFloras.org, Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
IPNI (2021). "Aster spathulatus Lindl". International Plant Names Index (IPNI). Royal Botanic Gardens, Kew; Harvard University Herbaria & Libraries; Australian National Botanic Gardens. Retrieved 22 July 2021.
NatureServe (2 July 2021a). "Symphyotrichum spathulatum Western Mountain Aster". explorer.natureserve.org. Arlington, Virginia. Retrieved 11 July 2021.
NatureServe (2 July 2021b). "Symphyotrichum spathulatum var. intermedium". explorer.natureserve.org. Arlington, Virginia. Retrieved 23 July 2021.
NatureServe (2 July 2021c). "Symphyotrichum spathulatum var. spathulatum". explorer.natureserve.org. Arlington, Virginia. Retrieved 23 July 2021.
NatureServe (2 July 2021d). "Symphyotrichum spathulatum var. yosemitanum". explorer.natureserve.org. Arlington, Virginia. Retrieved 23 July 2021.
POWO (2021a). "Symphyotrichum spathulatum (Lindl.) G.L.Nesom". Plants of the World Online. Royal Botanic Gardens, Kew. Retrieved 22 July 2021.
POWO (2021b). "Symphyotrichum spathulatum var. intermedium (A.Gray) G.L.Nesom". Plants of the World Online. Royal Botanic Gardens, Kew. Retrieved 22 July 2021.
POWO (2021c). "Symphyotrichum spathulatum var. yosemitanum (A.Gray) G.L.Nesom". Plants of the World Online. Royal Botanic Gardens, Kew. Retrieved 22 July 2021.
Semple, J.C. (n.d.a). "Symphyotrichum subsect. Occidentales Foliacei Asters". www.uwaterloo.ca. Ontario. Archived from the original on 28 June 2021. Retrieved 23 July 2021.
Semple, J.C. (n.d.b). "Symphyotrichum subg. Ascendentes". www.uwaterloo.ca. Ontario. Archived from the original on 22 July 2021. Retrieved 22 July 2021.
Semple, J.C. (17 May 2021a). "Classification of Symphyotrichum". www.uwaterloo.ca. Ontario. Archived from the original on 21 June 2021. Retrieved 23 July 2021.
Semple, J.C. (14 May 2021b). "Symphyotrichum ascendens — Long-leaved Aster, Intermountain Aster, Western Aster". www.uwaterloo.ca. Ontario. Archived from the original on 22 July 2021. Retrieved 22 July 2021.
Yatskievych, G., ed. (n.d.). "Symphyotrichum sect. Occidentales (Rydb.) G.L. Nesom". Flora of Missouri. Missouri Botanical Garden. Retrieved 23 July 2021 – via Tropicos.org.

[12] Occidentales is listed as a section in Flora of Missouri,^[10] whereas on Canadian botanist John C. Semple's Astereae Lab website, it is given as a subsection of section Symphyotrichum.^[11]

[FOOTNOTENatureServe2021a-1] NatureServe 2021a.

[FOOTNOTEPOWO2021a-2] POWO 2021a.

[FOOTNOTEPOWO2021b-3] POWO 2021b.

[FOOTNOTEPOWO2021c-4] POWO 2021c.

[FOOTNOTEBrouilletSempleAllenChambers2006a-5] Brouillet et al. 2006a.

[FOOTNOTESemplen.d.a-6] Semple n.d.a.

[FOOTNOTEBrouilletSempleAllenChambers2006c-7] Brouillet et al. 2006c.

[FOOTNOTEBrouilletSempleAllenChambers2006b-8] Brouillet et al. 2006b.

[FOOTNOTEAllen2012-9] Allen 2012.

[FOOTNOTEYatskievychn.d.-10] Yatskievych n.d.

[FOOTNOTESemple2021a-11] Semple 2021a.

[FOOTNOTEIPNI2021-13] IPNI 2021.

[FOOTNOTESemple2021b-14] Semple 2021b.

[FOOTNOTESemplen.d.b-15] Semple n.d.b.

[FOOTNOTEBrouilletSempleAllenChambers2006d-16] Brouillet et al. 2006d.

[FOOTNOTENatureServe2021b-17] NatureServe 2021b.

[FOOTNOTENatureServe2021c-18] NatureServe 2021c.

[FOOTNOTENatureServe2021d-19] NatureServe 2021d.

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PROFILPELAJAR.COM

Symphyotrichum spathulatum

Description

Chromosomes

Taxonomy

Distribution and habitat

Conservation

Notes

Citations

References

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