Gurlin Tsav skull

The "Gurlin Tsav" skull is a currently unnamed carnivorous metatherian fossil from the Nemegt Formation of Mongolia. Composed of a single semi-complete skull, this specimen is notable in regards to the evolution and systematics of Metatheria as a whole, and thus nigh-omnipresent in phylogenetic analyses of this group.^[1]^[2]^[3]^[4]

Fossil location

The Gurlin Tsav skull, as the informal name indicates, comes from the Gurlin Tsav track in the Nemegt Formation.^[5] This area represents fluvial deposits, and also heralds a few dinosaur specimens, such as a Saurolophus skeleton.^[6]

Classification

The specimen was originally referred to Deltatheroida, a clade of carnivorous metatherians rather common in the Cretaceous of Asia.^[7] However, one study by Guillermo Rougier has found it to lack the group's synapomorphies and instead resemble the American stagodontids more closely.^[2]

Posterior phylogenetic studies have indeed recovered this specimen as being closer to the Marsupialiformes line than to Deltatheroida. However, instead of being close to stagodontids, the Gurlin Tsav skull usually groups outside of a clade leading to South American metatherians (and, consequently, to marsupials). It is nearly always an immediate outgroup to sparassodonts.^[1]^[2]^[4]

Most recently, it has been found to group with sparassodonts and other Paleocene non-marsupial metatherians from South America. It and these taxa from a group independent from not only Deltatheroida, but also a North American clade where true marsupials are nested as well as stagodontids.^[4]

Marsupialiformes

Gurlin Tsav skull

Borhyaenidae

Mayulestes

Jaskhadelphys

	Andinodelphys

	Pucadelphys

Asiatherium

	Iugomortiferum

	Kokopellia

Aenigmadelphys

Anchistodelphys

Glasbiidae	Glasbius

Pediomyidae	Pediomys

Stagodontidae

Pariadens

	Eodelphis

	Didelphodon

Alphadontidae

	Turgidodon

	Alphadon

	Albertatherium

Marsupialia

References

^ ^a ^b S. Bi, X. Jin, S. Li and T. Du. 2015. A new Cretaceous metatherian mammal from Henan, China. PeerJ 3:e896
^ ^a ^b ^c Guillermo W. Rougier; Brian M. Davis; Michael J. Novacek (2015). "A deltatheroidan mammal from the Upper Cretaceous Baynshiree Formation, eastern Mongolia". Cretaceous Research. 52, Part A: 167–177. doi:10.1016/j.cretres.2014.09.009.
^ Eric J. Sargis, Marian Dagosto, Mammalian Evolutionary Morphology: A Tribute to Frederick S. Szalay, Springer Science & Business Media, 21/05/2008
^ ^a ^b ^c Wilson, G.P.; Ekdale, E.G.; Hoganson, J.W.; Calede, J.J.; Linden, A.V. (2016). "A large carnivorous mammal from the Late Cretaceous and the North American origin of marsupials". Nature Communications. 7: 13734. Bibcode:2016NatCo...713734W. doi:10.1038/ncomms13734. PMC 5155139. PMID 27929063.
^ "Fossilworks: Gateway to the Paleobiology Database".
^ S. Suzuki and M. Watabe. 2000. Report on the Japan–Mongolia Joint Paleontological Expedition to the Gobi desert, 1998. Hayashibara Museum of Natural Sciences Research Bulletin 1:83-98
^ Zofia Kielan-Jaworowska, Lev A. Nessov, On the metatherian nature of the Deltatheroida, a sister group of the Marsupialia, First published: January 1990Full publication history DOI: 10.1111/j.1502-3931.1990.tb01776.x