Brotia costula
preserved specimen
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Subclass:	Caenogastropoda
Superfamily:	Cerithioidea
Family:	Pachychilidae
Genus:	Brotia
Species:	B. costula
Binomial name
Brotia costula (Rafinesque, 1833)
Synonyms
see list

Brotia costula is a species of gastropod belonging to the family Pachychilidae.^[1][2]

This species is found in Southeastern Asia.^[1]

• Brotia (Brotia) costula (Rafinesque, 1833)
• Brotia (Brotia) costula costula (Rafinesque, 1833)
• Brotia (Brotia) costula varicosa (Troschel, 1837)
• Brotia variabilis (Benson, 1836) (based on an unavailable name)
• Melania (Melanoides) soloensis K. Martin, 1905 † (junior subjective synonym)
• Melania (Melanoides) variabilis Benson, 1836 (superseded combination)
• Melania (Melanoides) variabilis var. microstoma G. Nevill, 1885 (original combination)
• Melania (Melanoides) variabilis var. pseudospinosa G. Nevill, 1885 (superseded combination)
• Melania (Melanoides) variabilis var. semilaevigata G. Nevill, 1885 (original combination)
• Melania (Melanoides) variabilis var. subspinosa G. Nevill, 1885 (original combination)
• Melania (Melanoides) variabilis var. subtuberculata G. Nevill, 1885 (original combination)
• Melania (Melanoides) variabilis var. subvaricosa G. Nevill, 1885 (original combination)
• Melania carolinae Gray in Griffith & Pidgeon, 1833 (junior synonym)
• Melania costula Rafinesque, 1833 (original combination)
• Melania hainesiana I. Lea, 1857
• Melania indica Souleyet, 1852 (junior synonym)
• Melania menkeana "Brot, 1875" (incorrect subsequent spelling of Melania menkiana; not qualifying as an emendation)
• Melania menkeana G. Nevill, 1885 (unjustified emendation of Melania menkiana I. Lea, 1843)
• Melania menkiana I. Lea, 1843 (junior synonym)
• Melania meukiana I. Lea, 1843 (incorrect original spelling; Melania menkiana is an emendation)
• Melania plicata I. Lea, 1838 (invalid: junior homonym of Melania plicata Menke, 1828; M. menkiana is a replacement name)
• Melania spinosa Hanley, 1854 (junior synonym)
• Melania variabilis Benson, 1836 (invalid: junior homonym of Melania variabilis Defrance, 1823)
• Melania variabilis var. aspera Hanley & Theobald, 1874 (invalid: junior homonym of Melania aspera Lesson, 1831)
• Melania variabilis var. cincta Hanley & Theobald, 1874 (invalid: junior homonym of Melania cincta I. Lea & H. C. Lea, 1851)
• Melania variabilis var. microstoma G. Nevill, 1885 (invalid: junior homonym of Melania microstoma I. Lea & H. C. Lea, 1851)
• Melania variabilis var. pseudospinosa G. Nevill, 1885 (junior synonym)
• Melania variabilis var. semilaevigata G. Nevill, 1885 (junior synonym)
• Melania variabilis var. subspinosa G. Nevill, 1885 (junior synonym)
• Melania variabilis var. subtuberculata G. Nevill, 1885 (junior synonym)
• Melania variabilis var. subvaricosa G. Nevill, 1885 (junior synonym)
• Melania varicosa Troschel, 1837

• Köhler F. & Glaubrecht M. (2001) Toward a systematic revision of the southeast Asian freshwater gastropod Brotia H. Adams, 1866 (Cerithioidea: Pachychylidae): An account of species from around the South China Sea. Journal of Molluscan Studies 67: 281–318.

• Rafinesque, C. S. (1833). On 5 new fresh water shells, of Bengal and Assam in Asia. Atlantic Journal and Friend of Knowledge. 5: 165–170
• Benson, W. H. (1836). Descriptive catalogue of a collection of land and fresh-water shells, chiefly contained in the Museum of the Asiatic Society. Journal of the Asiatic Society of Bengal. 5(59): 741–750
• Griffith E. & Pidgeon E. (1833–1834). The Mollusca and Radiata. Vol. 12, In: E. Griffith, [1824−1835, The Animal Kingdom arranged in conformity with its organization, by the Baron Cuvier, London: Whittaker and Co., viii + 601 pp., 61 pls]
• Lea, I. (1838). Description of new freshwater and land shells. Transactions of the American Philosophical Society. new ser., 6: 1–154, plates 1–24
• Troschel, F. H. (1837). Neue Süßwasser-Conchylien aus dem Ganges. Archiv für Naturgeschichte. 3: 166–182
• Lea, I. (1857). Description of fifteen new species of exotic Melaniana. Proceedings of the Academy of Natural Sciences of Philadelphia. 8(4): 144–145
• Lea, 1843) Lea, I. (1843). On fresh water shells (continuation) [sixteen new species of the family Melanianae. Proceedings of the American Philosophical Society. 2(24): 241–243]
• Nevill G. (1885). Hand List of Mollusca in the Indian Museum, Calcutta. Part II. Gastropoda. Prosobranchia-Neurobranchia (contd.). x + 306 pp. Office of Superintendent of Government Printing, Calcutta
• Martin K. (1895–1906). Die Fossilien von Java auf grund einer Sammlung von Dr. R. D. M. Verbeek. I. Band. Gasteropoda. Sammlungen des Geologischen Reichs-Museums in Leiden. new ser., 1: 1–132, pls 1–20
• Köhler, F.; Glaubrecht, M. (2006). A systematic revision of the southeast Asian freshwater gastropod Brotia (Cerithioidea: Pachychilidae). Malacologia. 48(1–2): 159–251