Vikinghøgda Formation

Vikinghøgda Formation
Stratigraphic range: Induan-Olenekian (Griesbachian-Spathian)
~252–247.2 Ma
TypeFormation
Unit ofSassendalen Group
Sub-unitsDeltadalen Member, Lusitaniadalen Member, Vendomdalen Member
UnderliesBotneheia Formation
OverliesKapp Starostin Formation
Lithology
PrimaryMudstone, siltstone
OtherShale, sandstone
Location
Approximate paleocoordinates42°12′N 10°24′E / 42.2°N 10.4°E / 42.2; 10.4
RegionSvalbard
CountryNorway
Vikinghøgda Formation is located in Svalbard
Vikinghøgda Formation
Vikinghøgda Formation (Svalbard)

The Vikinghøgda Formation is a geologic formation in Svalbard, Norway. It preserves fossils dating back to the Early Triassic (Griesbachian-Spathian) period.[1] It is split into three members, from oldest to youngest: the Deltadalen Member (Induan), Lusitaniadalen Member (Smithian), and Vendomdalen Member (Spathian). The formation can be found in central Spitsbergen (Sassendalen, Dickson Land, James I Land), southern Spitsbergen, as well as the smaller islands of Barentsøya and Edgeøya. The type locality is positioned in the vicinity of Vikinghøgda and Sticky Keep, two low peaks along the southeast edge of Sassendalen (Sassen Valley) in Spitsbergen. The two upper members of the Vikinghøgda Formation were previously grouped together as the Sticky Keep Formation.[2][3]

Subunits

The Vikinghøgda Formation is formed by fine-grained marine sediments, such as mudstones, shales, siltstones, and fine sandstones. There is a trend of finer sediments, deeper waters, and higher organic content through the formation. Though fossils are only abundant in the middle part (Lusitaniadalen Member) of the formation, index fossils can be found throughout the whole formation. The Vikinghøgda Formation is one of the better records of Early Triassic chronostratigraphy in the Boreal realm, owing to a combination of continuous sedimentation, distinctive index fossils, palynomorph stratigraphy, magnetostratigraphy,[2][4] and trace metal cyclostratigraphy.[5]

Deltadalen Member

The Deltadalen Member is named after the small stream valley which runs between Vikinghøgda and Sticky Keep. The member consists of about 70 meters of silty shales and fine sandstones, lying above the eroded surface of the Permian-age Kapp Starostin Formation. Some of the sandstone beds may be glauconitic or hummocky, and calcareous nodules may be present.[2] Fossils are rare and restricted to these nodules, though they include a variety of silicified ammonoids, conodonts, bivalves, and gastropods.[6] Some index fossils have biostratigraphic significance, such as the ammonoid Otoceras boreale and the conodonts Neogondolella carinata and Neospathodus svalbardensis. These species constrain the member to the Induan stage, between the early Griesbachian and early Dienerian substages. The sandiest intervals can be found at the base and the top of the member. The depositional environment is reconstructed as a shallow marine setting influenced by storms and nearby deltaic sediments.[2]

The Deltadalen Member has been equated with two other Induan-age formations in its vicinity: the Vardebukta Formation (exposed in western and southern Spitsbergen) and the Havert Formation (under the Barents Sea).[2]

A 2020 study reported that the Permian-Triassic boundary occurs near the start of the Deltadalen Member. This was supported by several lines of evidence. The conodont Hindeodus parvus, which defines the base of the Triassic, was reported from a sediment layer 4.1 meters above the base of the member. An overlying tephra bed was dated to 252.13 ± 0.62 Ma via U-Pb radiometric dating, an age which is congruent with other reported estimates for the boundary.[5]

Lusitaniadalen Member

The Lusitaniadalen Member is named after a glacier on the western flank of Vikinghøgda. The member is about 90 meters thick, mostly composed of laminated silty mudstones. Sandstone beds are less common, though when they occur, they are hummocky. Fossiliferous calcareous nodules are abundant in the sandstone-bearing layers, encasing well-preserved fossils of both invertebrates and vertebrates. The ammonoid index fossils Euflemingites romunderi and Wasatchites tardus indicate that the Lusitaniadalen Member was deposited during the Smithian substage of the Olenekian stage. The most common ammonoid is Arctoceras.[7] Fish and amphibian fossils have also been found in this member, particularly at the "Fish Niveau" bonebed. This horizon also produced the oldest known ichthyopterygian reptile fossil.[8] The depositional environment corresponds to a deeper and calmer conditions, indicative of a major transgression (sea level rise) affecting the continental shelf.[2]

The Lusitaniadalen Member has been equated with the Iskletten Member of the Tvillingodden Formation (in western Spitsbergen) and the lower part of the Klappmyss Formation (under the Barents Sea). It has also been previously described as the Iskletten Member of the Sticky Keep Formation.[2]

Vendomdalen Member

Dark shale (laminated mudstone) dominates the 90-meter-thick Vendomdalen Member, though silty yellowish dolomitic beds also occur. Very large dolomite nodules or lenses, some over a meter in width, can also be found in this member. However, fossil-bearing concretions are rarer and more deformed than in the Lusitaniadalen Member. Fossils include abundant bivalves and Spathian ammonoid index fossils, such as Bajarunia euomphala, Keyserlingites subrobustus, and Parasibirites elegans.[2] Marine reptile and fish bonebeds also occur in the Vendomdalen Member, the two most prominent being the older "Grippia Niveau" and younger "Lower Saurian Niveau".[9][10][11][12] The Vendomdalen Member corresponds to the deepest part of the continental shelf, with high organic matter deposition and no influence from storm events.[2]

The Vendomdalen Member has been equated with the Kaosfjellet Member of the Tvillingodden Formation (in western Spitsbergen) and the upper part of the Klappmyss Formation (under the Barents Sea). It has also been previously described as the Kaosfjellet Member of the Sticky Keep Formation.[2]

Paleobiota

Tetrapods

Tetrapod remains occur in several horizons of the Vikinghøgda Formation. The fossils belong to ichthyosauriform and possibly sauropterygian reptiles and temnospondyl amphibians.

Reptiles

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
Reptiles of the Vikinghøgda Formation
Taxon / Genus Species Subunit Notes Images
Cymbospondylus C. sp. Vendomdalen Member (Lower Saurian Niveau) A large basal ichthyosaur[9][12]
Grippia longirostris
Reptilia indet. Vendomdalen Member (Grippia Niveau) An indeterminate reptile fossil (SVT 203) previously referred to Grippia longirostris. Postcranial proportions differ from all known ichthyosauromorphs, and may instead suggest affinities with Helveticosaurus or placodonts.[13]
Grippia G. longirostris Vendomdalen Member (Grippia Niveau) A small basal ichthyopterygian[9]
Ichthyopterygia indet. Lusitaniadalen Member (Fish Niveau),[8] Vendomdalen Member[14] Indeterminate ichthyopterygian remains. Pelagic-adapted ichthyopterygian fossils from the Lusitaniadalen Member are the oldest known from Svalbard and the world.[8]
Isfjordosaurus I. minor Vendomdalen Member (Lower Saurian Niveau) An enigmatic ichthyosauromorph, possibly a hupehsuchian or basal ichthyopterygian[9]
Merriamosaurus M. hulkei Vendomdalen Member (Lower Saurian Niveau) An ichthyopterygian based on forelimb fossils, initially described under the preoccupied name Rotundopteryx hulkei. Most likely a junior synonym of Pessopteryx nisseri.[9]
Omphalosaurus O. nisseri Vendomdalen Member (Lower Saurian Niveau) Weathered jaw fragments and other omphalosaurid-like fossils of uncertain validity, some of which have also been named as O. merriami or referred to Pessopteryx nisseri.[9]
O. sp.[14] Vendomdalen Member (Grippia and Lower Saurian Niveaus) Some fossils referred to Omphalosaurus appear to legitimately belong to the genus.[14]
Pessopteryx P. nisseri Vendomdalen Member (Lower Saurian Niveau) A large shastasaurid-like ichthyosaur, previously utilized as a chimera encompassing various ichthyopterygian fossils from the Lower Saurian Niveau.[9][12]
Quasianosteosaurus Q. vikinghoegdai Vendomdalen Member (Grippia Niveau) A medium-sized ichthyopterygian[9]
Svalbardosaurus S. crassidens Vendomdalen Member (Grippia Niveau) A dubious tetrapod based on large teeth initially referred to ichthyosaurs, but more likely belonging to temnospondyl amphibians.[9]

Amphibians

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
Temnospondyls of the Vikinghøgda Formation
Taxon / Genus Species Subunit Notes Images
Aphaneramma A. rostratum Lusitaniadalen Member (Fish Niveau) A long-snouted lonchorhynchine trematosaurid[15]
Aphaneramma
Boreosaurus B. thorslundi Lusitaniadalen Member (Fish Niveau) A brachyopid or rhytidosteid[15]
Lyrocephaliscus L. euri Lusitaniadalen Member (Fish Niveau) A trematosaurine trematosaurid, formerly known as Lyrocephalus.[15]
Peltostega P. erici Lusitaniadalen Member (Fish Niveau) A rhytidosteid[15]
Platystega P. depressa Lusitaniadalen Member (Fish Niveau) A trematosaurine trematosaurid[15]
Tertrema T. acuta Lusitaniadalen Member (Fish Niveau) A relatively short-snouted trematosaurid, possibly a lonchorhynchine[16]
Sassenisaurus S. spitzbergensis Lusitaniadalen Member (Fish Niveau) Possibly an early capitosaur[15]

Fish

A diverse fish assemblage has been described from the late Smithian[17] (early Olenekian) aged "Fish Niveau" (Lusitaniadalen Member, lower part of the "Sticky Keep Member") and other horizons, including lungfish, coelacanths, ray-finned fish, and hybodont sharks.[18][11][19] Most taxa were described by Erik Stensiö.[20][21] The lists below are based on Kogan & Romano[17] and Bratvold et al.[11]

Lobe-finned fish

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
Sarcopterygii of the Vikinghøgda Formation
Taxon / Genus Species Subunit Notes Images
Ceratodus C. sp. Vendomdalen Member (below Lower Saurian Niveau) Tooth plate of a lungfish[20]
Ceratodus
Axelia[20] A. robusta Lusitaniadalen Member (Fish Niveau) A coelacanthid coelacanth
A. elegans Lusitaniadalen Member (Fish Niveau) A coelacanthid coelacanth
Mylacanthus[20] M. lobatus Lusitaniadalen Member (Fish Niveau) A coelacanthid coelacanth, quite similar to Axelia.[22]
M. spinosus Lusitaniadalen Member (Fish Niveau) A coelacanthid coelacanth, quite similar to Axelia.[22]
Sassenia[20] S. tuberculata Lusitaniadalen Member (Fish Niveau) A rhabdodermatid coelacanth
S.? guttata Lusitaniadalen Member (Fish Niveau) A rhabdodermatid coelacanth? Dubious generic attribution.[22]
S. sp. Lusitaniadalen Member (Fish Niveau) A rhabdodermatid coelacanth
Scleracanthus[20] S. asper Lusitaniadalen Member (Fish Niveau) A coelacanth, quite similar to Axelia.[22]
Wimania[20] W. sinuosa Lusitaniadalen Member (Fish Niveau) A relatively large coelacanthid coelacanth
W.? multistriata Lusitaniadalen Member (Fish Niveau) A coelacanth
W.? sp. Lusitaniadalen Member (Fish Niveau) A coelacanth

Ray-finned fish

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
Actinopterygii of the Vikinghøgda Formation
Taxon / Genus Species Subunit Notes Images
Acrorhabdus[20] A. bertili Lusitaniadalen Member (Fish Niveau) A ptycholepid
Saurichthys
A. asplundi Lusitaniadalen Member (Fish Niveau) A ptycholepid
A. latistriatus Lusitaniadalen Member (Fish Niveau) A ptycholepid
Birgeria B. aldingeri Lusitaniadalen Member (Fish Niveau) A birgeriid.The material was originally referred to B. mougeoti,[20] the type species of Birgeria, but was later ascribed to its own species
B. sp. Lusitaniadalen Member (Fish Niveau) A partial skull of a large Birgeria[23]
Bobasatrania B. nathorsti Lusitaniadalen Member (Fish Niveau) A large bobasatraniform that was originally referred to as Platysomus nathorsti[20]
Boreosomus B. arcticus Lusitaniadalen Member (Fish Niveau) A ptycholepid and the type species of Boreosomus, which was originally described as Acrolepis arctica by Arthur Smith Woodward
B. reuterskioldi Lusitaniadalen Member (Fish Niveau) A ptycholepid[20]
B.? scaber[20] Lusitaniadalen Member (Fish Niveau)
Pteronisculus P. gyrolepidoides Lusitaniadalen Member (Fish Niveau) A poorly known rhadinichthyid or turseoid, It was originally described as Glaucolepis gyrolepidoides,[20] but later referred to the genus Pteronisculus because the genus name Glaucolepis is preoccupied
Pygopterus[20] P. degeeri Lusitaniadalen Member (Fish Niveau) A pygopterid. Pygopterus is one of the genera of ray-finned fish that crossed the Permian-Triassic boundary
Saurichthys S. elongatus Lusitaniadalen Member (Fish Niveau) and possibly in beds slightly above it A saurichthyiform with a slender, elongated rostrum[21]
S. hamiltoni Lusitaniadalen Member (Fish Niveau) A saurichthyiform[21]
S. ornatus Lusitaniadalen Member (Fish Niveau) A saurichthyiform. This species is known primarily from skull remains[21][17]
S. ornatus? Lusitaniadalen Member (Fish Niveau) A saurichthyiform. Based on one specimen that shows the postcranium and which has a different scale pattern than S. ornatus[17]
S. wimani Lusitaniadalen Member (Fish Niveau) A saurichthyiform. Based on a specimen preserving the skull and anteriormost body portion that was originally described as "Belonorhynchus" wimani by Arthur Smith Woodward. A later found postcranium was referred to this species[17]
S. sp. Lusitaniadalen Member (Fish Niveau) A saurichthyiform[21]
Teffichthys T. woodwardi Lusitaniadalen Member (Fish Niveau) A perleidiform. It was originally described as Colobodus altilepis by Arthur Smith Woodward but later referred to as Perleidus woodwardi[20] (the species name was changed to avoid confusion with the type species of Perleidus, P. altolepis). The Early Triassic species of Perleidus were later ascribed to a different genus, Teffichthys[24]

Cartilaginous fish

Chondrichthyan remains are predominantely known from the Vendomdalen Member (Grippia Niveau). Several species listed below are also present in the nearby Vardebukta Formation, of Dienerian (late Induan) age.

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
Chondrichthyes of the Vikinghøgda Formation
Taxon / Genus Species Subunit Notes Images
Acrodus[20] A. gaillardoti[11] Vendomdalen Member (Grippia Niveau) An acrodontid hybodontiform shark. Only known from isolated teeth
Hybodus
A. lateralis[11] Vendomdalen Member (Grippia Niveau) An acrodontid hybodontiform shark. Only known from isolated teeth
A. oppenheimeri Vendomdalen Member (Grippia Niveau) An acrodontid hybodontiform shark. Only known from isolated teeth[11]
A. scaber Vendomdalen Member (Grippia Niveau) An acrodontid hybodontiform shark. Only known from isolated teeth[11]
A. spitzbergensis Vendomdalen Member (Grippia Niveau) An acrodontid hybodontiform shark. Only known from isolated teeth[11]
A. vermiformis Vendomdalen Member (Grippia Niveau) An acrodontid hybodontiform shark. Only known from isolated teeth[11]
A. sp.[11] Vendomdalen Member (Grippia Niveau) An acrodontid hybodontiform shark. Only known from isolated teeth
Hybodus[20] H. microdus/
H. ?microdus[11]
Vendomdalen Member (Grippia Niveau) A hybodontid hybodontiform shark. Only known from isolated teeth
H. rapax[11] Vendomdalen Member (Grippia Niveau) A hybodontid hybodontiform shark. Only known from isolated teeth
H. sasseniensis[11] Vendomdalen Member (Grippia Niveau) A hybodontid hybodontiform shark. Only known from isolated teeth
Lissodus L. angulatus Vendomdalen Member (Grippia Niveau) A lonchidiid hybodontiform shark. Only known from isolated teeth that were originally described as Polyacrodus angulatus
Nemacanthus N. sp. Vendomdalen Member (Grippia Niveau) A palaeospinacid synechodontiform shark. Only known from fin spines[11]
Palaeobates P. polaris Lusitaniadalen Member (Fish Niveau),
Vendomdalen Member (Grippia Niveau)
An acrodontid hybodontiform shark known from teeth[11] and skelettal elements. Skull and postcranial elements are only known from the Fish Niveau[20][25]
P. sp. Vendomdalen Member (Grippia Niveau) An acrodontid hybodontiform shark. Only known from isolated teeth[20]
Polyacrodus P. sp. Vendomdalen Member (Grippia Niveau) A polyacrodontid hybodontiform shark known from skull[11]
Hybodontoidea Gen. et sp. indet.1 Vendomdalen Member (Grippia Niveau) Basal plate of cephalic spine[11]
Gen. et sp. indet.2 Vendomdalen Member (Grippia Niveau) Fin spine[11]
Gen. et sp. indet.3 Vendomdalen Member (Grippia Niveau) Fin spine[11]
Synechodontiformes Gen. et sp. indet.1 Vendomdalen Member (Grippia Niveau) Isolated teeth[11]
Gen. et sp. indet.2 Vendomdalen Member (Grippia Niveau) Isolated teeth[11]

See also

References

  1. ^ Vikinghøgda Formation at Fossilworks.org
  2. ^ a b c d e f g h i j Mørk, Atle; Elvebakk, Geir; Forsberg, Arne W.; Hounslow., Mark W.; Nakrem., Hans Arne; Vigran, Jorunn Os; Weitschat, Wolfgang (1999). "The type section of the Vikinghøgda Formation: a new Lower Triassic unit in central and eastern Svalbard". Polar Research. 18 (1): 51–82. Bibcode:1999PolRe..18...51M. doi:10.1111/j.1751-8369.1999.tb00277.x (inactive 1 November 2024). ISSN 0800-0395.{{cite journal}}: CS1 maint: DOI inactive as of November 2024 (link)
  3. ^ Hurum, Jørn Harald; Engelschiøn, Victoria S.; Økland, Inghild; Bratvold, Janne; Ekeheien, Christina P.; Roberts, Aubrey J.; Delsett, Lene Liebe; Hansen, Bitten B.; Mørk, Atle; Nakrem, Hans Arne; Druckenmiller, Patrick S.; Hammer, Øyvind (2018-10-04). "The history of exploration and stratigraphy of the Early to Middle Triassic vertebrate-bearing strata of Svalbard (Sassendalen Group, Spitsbergen)". Norwegian Journal of Geology. 98 (2): 165–174. doi:10.17850/njg98-2-04. hdl:10852/71101. S2CID 216704366.
  4. ^ Hounslow, M. W.; Peters, C.; Mork, A.; Weitschat, W.; Vigran, J. O. (2008-09-01). "Biomagnetostratigraphy of the Vikinghogda Formation, Svalbard (Arctic Norway), and the geomagnetic polarity timescale for the Lower Triassic". Geological Society of America Bulletin. 120 (9–10): 1305–1325. Bibcode:2008GSAB..120.1305H. doi:10.1130/B26103.1. ISSN 0016-7606.
  5. ^ a b Zuchuat, V.; Sleveland, A. R. N.; Twitchett, R. J.; Svensen, H. H.; Turner, H.; Augland, L. E.; Jones, M. T.; Hammer, Ø.; Hauksson, B. T.; Haflidason, H.; Midtkandal, I.; Planke, S. (2020-09-15). "A new high-resolution stratigraphic and palaeoenvironmental record spanning the End-Permian Mass Extinction and its aftermath in central Spitsbergen, Svalbard". Palaeogeography, Palaeoclimatology, Palaeoecology. 554: 109732. Bibcode:2020PPP...55409732Z. doi:10.1016/j.palaeo.2020.109732. hdl:10852/81143. ISSN 0031-0182. S2CID 218955779.
  6. ^ Foster, William J.; Danise, Silvia; Twitchett, Richard J. (2017-10-03). "A silicified Early Triassic marine assemblage from Svalbard". Journal of Systematic Palaeontology. 15 (10): 851–877. doi:10.1080/14772019.2016.1245680. hdl:2158/1152798. ISSN 1477-2019. S2CID 54030401.
  7. ^ Hansen, Bitten Bolvig; Bucher, Hugo; Schneebeli-Hermann, Elke; Hammer, Øyvind (2021). "The middle Smithian (Early Triassic) ammonoid Arctoceras blomstrandi: conch morphology and ornamentation in relation to stratigraphy". Papers in Palaeontology. 7 (3): 1435–1457. Bibcode:2021PPal....7.1435H. doi:10.1002/spp2.1348.
  8. ^ a b c Kear, Benjamin P.; Engelschiøn, Victoria S.; Hammer, Øyvind; Roberts, Aubrey J.; Hurum, Jørn H. (2023). "Earliest Triassic ichthyosaur fossils push back oceanic reptile origins". Current Biology. 33 (5): R178–R179. doi:10.1016/j.cub.2022.12.053. PMID 36917937. S2CID 257498390.
  9. ^ a b c d e f g h i Maxwell, Erin E.; Kear, Benjamin P. (2013). "Triassic ichthyopterygian assemblages of the Svalbard archipelago: a reassessment of taxonomy and distribution". GFF. 135 (1): 85–94. Bibcode:2013GFF...135...85M. doi:10.1080/11035897.2012.759145. ISSN 1103-5897. S2CID 129092001.
  10. ^ Hansen, Bitten Bolvig; Hammer, Øyvind; Nakrem, Hans Arne (2018-10-04). "Stratigraphy and age of the Grippia niveau bonebed, Lower Triassic Vikinghøgda Formation, Spitsbergen" (PDF). Norwegian Journal of Geology. 98 (2): 175–187. doi:10.17850/njg98-2-02. S2CID 134204517.
  11. ^ a b c d e f g h i j k l m n o p q r s t u Bratvold, Janne; Delsett, Lene Liebe; Hurum, Jørn Harald (2018-10-04). "Chondrichthyans from the Grippia bonebed (Early Triassic) of Marmierfjellet, Spitsbergen". Norwegian Journal of Geology. 98 (2): 189–217. doi:10.17850/njg98-2-03. hdl:10852/71103. S2CID 132293043.
  12. ^ a b c Engelschiøn, Victoria Sjøholt; Delsett, Lene Liebe; Roberts, Aubrey Jane; Hurum, Jørn Harald (2018-10-04). "Large-sized ichthyosaurs from the Lower Saurian niveau of the Vikinghøgda Formation (Early Triassic), Marmierfjellet, Spitsbergen". Norwegian Journal of Geology. 98 (2): 239–265. doi:10.17850/njg98-2-05. hdl:10852/71102. S2CID 135275680.
  13. ^ Motani, Ryosuke (2000). "Skull of Grippia longirostris : no contradiction with a diapsid affinity for the Ichthyopterygia". Palaeontology. 43 (1): 01–14. Bibcode:2000Palgy..43....1M. doi:10.1111/1475-4983.00115. S2CID 129452053.
  14. ^ a b c Ekeheien, Christina Pokriefke; Delsett, Lene Liebe; Roberts, Aubrey Jane; Hurum, Jørn Harald (2018-10-04). "Preliminary report on ichthyopterygian elements from the Early Triassic (Spathian) of Spitsbergen". Norwegian Journal of Geology. 98 (2): 219–238. doi:10.17850/njg98-2-07. hdl:10852/71099. S2CID 134421320.
  15. ^ a b c d e f Kear, Benjamin P.; Poropat, Stephen F.; Bazzi, Mohamad (2016). "Late Triassic capitosaurian remains from Svalbard and the palaeobiogeographical context of Scandinavian Arctic temnospondyls". Geological Society, London, Special Publications. 434 (1): 113–126. Bibcode:2016GSLSP.434..113K. doi:10.1144/SP434.11. ISSN 0305-8719. S2CID 131531829.
  16. ^ Slodownik, Miriam A.; Mörs, Thomas; Kear, Benjamin P. (2021). "Reassessment of the Early Triassic trematosaurid temnospondyl Tertrema acuta from the Arctic island of Spitsbergen". Journal of Vertebrate Paleontology. 41 (1): e1900209. Bibcode:2021JVPal..41E0209S. doi:10.1080/02724634.2021.1900209. ISSN 0272-4634. S2CID 235574341.
  17. ^ a b c d e Kogan, Ilja; Romano, Carlo (2016). "A new postcranium of Saurichthys from the Early Triassic of Spitsbergen" (PDF). Freiberger Forschungshefte C (Paläontologie, Stratigraphie, Fazies 23). 550: 205–221. ISBN 9783860125526.
  18. ^ Cox, C. B.; Smith, David G. (1973). "A review of the Triassic vertebrate faunas of Svalbard". Geological Magazine. 110 (5): 405–418. Bibcode:1973GeoM..110..405C. doi:10.1017/S0016756800036190. ISSN 0016-7568. S2CID 128809142.
  19. ^ Brinkmann, Winand; Romano, Carlo; Bucher, Hugo; Ware, David; Jenks, Jim (2010). "Palaeobiogeography and stratigraphy of advanced gnathostomian fishes (Chondrichthyes and Osteichthyes) in the Early Triassic and from selected Anisian localities (Report 1863–2009)" (PDF). Zentralblatt für Geologie und Paläontologie, Teil II. 2009: 765–812..
  20. ^ a b c d e f g h i j k l m n o p q r s Stensiö, E. (1921). Triassic fishes from Spitzbergen 1. Wien: Adolf Holzhausen. pp. xxviii+307.
  21. ^ a b c d e Stensiö, E. (1925). "Triassic fishes from Spitzbergen 2". Kungliga Svenska Vetenskapsakademiens Handlingar. 3: 1–261.
  22. ^ a b c d Forey, Peter L. (1998). History of the coelacanth fishes. London: Chapman & Hall. p. 434. ISBN 9780412784804.
  23. ^ Scheyer, Torsten M.; Romano, Carlo; Jenks, Jim; Bucher, Hugo (2014). "Early Triassic Marine Biotic Recovery: The Predators' Perspective". PLOS ONE. 9 (3): e88987. Bibcode:2014PLoSO...988987S. doi:10.1371/journal.pone.0088987. PMC 3960099. PMID 24647136.
  24. ^ Marramà, G.; Lombardo, C.; Tintori, A. & Carnevale, G. (2017). "Redescription of "Perleidus"(Osteichthyes, Actinopterygii) from the Early Triassic of northwestern Madagascar". Rivista Italiana di Paleontologia e Stratigrafia. 123 (2): 219–242. doi:10.13130/2039-4942/8328.
  25. ^ Romano, Carlo; Brinkmann, Winand (December 2010). "A new specimen of the hybodont shark Palaeobates polaris with threedimensionally preserved Meckel's cartilage from the Smithian (Early Triassic) of Spitsbergen". Journal of Vertebrate Paleontology. 30 (6): 1673–1683. Bibcode:2010JVPal..30.1673R. doi:10.1080/02724634.2010.521962. S2CID 86411191.

Read other articles:

Anton de KomLahirCornellis Gerrad Anton de Kom22 Februari 1898Paramaribo, SurinameMeninggal24 April 1945Sandbostel, JermanKarya terkenalPenulis Buku Wij Slaven Van Suriname (We Slaves Of Suriname)Suami/istriPetronella BorsboomAnakCees De KomOrang tuaAdolf De Kom dan Judith Jacoba Dulder Patung Anton de Kom Anton De Kom nama lengkap Cornellis Gerrad Anton De Kom (22 Februari 1898 – 24 April 1945) adalah Penulis, Militer dan Aktivis Suriname. Namanya diabadikan sebagai nama Uni...

 

 

Untuk nama novel dengan judul yang sama seperti film ini, lihat 5 cm. 5 cmPoster filmSutradaraRizal MantovaniProduserSunil SorayaSkenarioDonny DhirgantoroSunil SorayaHilman MutasiBerdasarkan5 cmoleh Donny DhirgantoroPemeranHerjunot AliRaline ShahFedi NurilPevita PearceDenny SumargoSaykojiPenata musikNidjiSinematograferYudi DatauPenyuntingSastha SunuPerusahaanproduksiSoraya Intercine FilmsTanggal rilis12 Desember 2012 (2012-12-12)Durasi125 menitNegara IndonesiaBahasaBahasa Indonesia...

 

 

Song by Big K.R.I.T. Country ShitSingle by Big K.R.I.T.from the album K.R.I.T. Wuz Here ReleasedSeptember 28, 2010 (2010-09-28)Recorded2010GenreHip hop, Southern hip hopLength3:15LabelIsland Def JamSongwriter(s)Christopher Bridges, Bernard Freeman, Kenneth Gamble, Leon Huff, Justin L ScottBig K.R.I.T. singles chronology Country Shit (2010) Moon & Stars (Remix) (2011) Country Shit is a song by rapper Big K.R.I.T. The remix features rappers Ludacris and Bun B. The song wa...

2002 Canadian filmEvelyn: The Cutest Evil Dead GirlDirected byBrad PeytonWritten byBrad PeytonLee HoverdProduced byJames MauroStarringNadia LitzJoshua CloseCinematographyDylan MacLeodEdited byKathy WeinkaufMusic byPhilip StangerRelease date 2002 (2002) Running time9 minutesCountryCanadaLanguageEnglishBudget$16,000 Evelyn: The Cutest Evil Dead Girl is a 2002 black comedy short film[1] directed by Newfoundlander Brad Peyton. It stars Nadia Litz in the title role, and Joshua Close, ...

 

 

Pour les articles homonymes, voir Aragua. Cet article est une ébauche concernant le Venezuela. Vous pouvez partager vos connaissances en l’améliorant (comment ?) selon les recommandations des projets correspondants. État d'Aragua Héraldique Drapeau La ville de Maracay, capitale de l'État d'Aragua, est l'une des cinq principales agglomérations urbaines du Venezuela. Administration Pays Venezuela Capitale Maracay Gouverneur Mandat Karina Carpio Depuis 2021 (en cours) Démographie ...

 

 

Chronologies Données clés 1576 1577 1578  1579  1580 1581 1582Décennies :1540 1550 1560  1570  1580 1590 1600Siècles :XIVe XVe  XVIe  XVIIe XVIIIeMillénaires :-Ier Ier  IIe  IIIe Chronologies thématiques Art Architecture, Arts plastiques (Dessin, Gravure, Peinture et Sculpture), (), Littérature () et Musique (Classique)   Ingénierie (), Architecture et ()   Politique Droit   Religion (,)   Science () et Santé...

У этого термина существуют и другие значения, см. Собор Александра Невского. Православный храмКафедральный собор Святого Александра НевскогоCathédrale Saint-Alexandre-Nevsky 48°52′39″ с. ш. 2°18′07″ в. д.HGЯO Страна  Франция Город Париж, улица Дарю, 12 Конфессия Православие Е�...

 

 

この記事は検証可能な参考文献や出典が全く示されていないか、不十分です。出典を追加して記事の信頼性向上にご協力ください。(このテンプレートの使い方)出典検索?: コルク – ニュース · 書籍 · スカラー · CiNii · J-STAGE · NDL · dlib.jp · ジャパンサーチ · TWL(2017年4月) コルクを打ち抜いて作った瓶の栓 コルク(木栓、�...

 

 

Questa voce sugli argomenti allenatori di pallacanestro turchi e cestisti turchi è solo un abbozzo. Contribuisci a migliorarla secondo le convenzioni di Wikipedia. Segui i suggerimenti del progetto di riferimento. Orhun Ene Orhun Ene nel 2018 Nazionalità  Turchia Altezza 188 cm Pallacanestro Ruolo Allenatore (ex playmaker) Squadra  Tofaş Bursa Termine carriera 2003 - giocatore CarrieraSquadre di club 1978-1981Ortakoyspor1981-1989 Eczacıbaşı1989-1992 Paşabahçe...

此條目可参照英語維基百科相應條目来扩充。 (2021年5月6日)若您熟悉来源语言和主题,请协助参考外语维基百科扩充条目。请勿直接提交机械翻译,也不要翻译不可靠、低品质内容。依版权协议,译文需在编辑摘要注明来源,或于讨论页顶部标记{{Translated page}}标签。 约翰斯顿环礁Kalama Atoll 美國本土外小島嶼 Johnston Atoll 旗幟颂歌:《星條旗》The Star-Spangled Banner約翰斯頓環礁�...

 

 

Irish-Welsh cyclist Matt BrammeierBrammeier at the 2015 Scheldeprijs.Personal informationFull nameMatthew Martin BrammeierBorn (1985-06-07) 7 June 1985 (age 38)Liverpool, EnglandTeam informationCurrent teamRetiredDisciplinesRoadTrackRoleRider (retired)CoachProfessional teams2006DFL-Cyclingnews-Litespeed2007–2008Profel Ziegler Continental Team2009–2010An Post–Sean Kelly2011HTC–Highroad2012Omega Pharma–Quick-Step[1]2013Champion System2014Synergy Baku[2]2...

 

 

此条目序言章节没有充分总结全文内容要点。 (2019年3月21日)请考虑扩充序言,清晰概述条目所有重點。请在条目的讨论页讨论此问题。 哈萨克斯坦總統哈薩克總統旗現任Қасым-Жомарт Кемелұлы Тоқаев卡瑟姆若马尔特·托卡耶夫自2019年3月20日在任任期7年首任努尔苏丹·纳扎尔巴耶夫设立1990年4月24日(哈薩克蘇維埃社會主義共和國總統) 哈萨克斯坦 哈萨克斯坦政府...

Zachary Taylor's plantation, Jefferson Co. Mississippi Map from 1866 showing Genl. Taylors Old Place The Cypress Grove Plantation was a Southern plantation owned by President Zachary Taylor near Rodney, Mississippi. Later, it was also known as Buena Vista Plantation.[1] Location The plantation bordered Ashland Plantation and the Mississippi River.[1] It was near the Ashland community, and Ashland's port was sometimes used by Cypress Grove Plantation.[2] The plantation ...

 

 

PT Balai PustakaPerusahaan indukDanareksaStatusBeroperasiDidirikan22 September 1917; 106 tahun lalu (1917-09-22)PendiriAdviseur voor Inlandsch ZakenNegara asalIndonesiaKantor pusatJakartaTokoh kunciAchmad Fachrodji (Direktur Utama)Jenis terbitanBuku sastraBuku pendidikanBuku bacaan umumKamusPustaka DigitalTaman Bacaan MasyarakatSitus resmiwww.balaipustaka.co.id Balai Pustaka (Ejaan Van Ophuijsen: Balai Poestaka) adalah perusahaan yang bergerak di bidang penerbitan, percetakan dan multime...

 

 

Parlemen Antigua dan Barbuda Parliament of Antigua and BarbudaJenisJenisBicameral MajelisDewan Perwakilan RakyatSenatAnggota34 orang17 anggota Dewan17 anggota SenatPemilihanSistem pemilihan Dewan Perwakilan RakyatFirst-past-the-postSistem pemilihan SenatPengangkatan oleh Gubernur JenderalTempat bersidangSt. John'sSitus webwww.ab.gov.ag/gov_v1/parliament/index.htm L • BBantuan penggunaan templat ini Parlemen Antigua dan Barbuda adalah lembaga legislatif nasional Antigua dan Barbuda. Lem...

My Night at Maud'sSampul DVDSutradaraÉric RohmerProduserPierre CottrellBarbet SchroederDitulis olehÉric RohmerPemeranJean-Louis TrintignantFrançoise FabianMarie-Christine BarraultAntoine VitezSinematograferNéstor AlmendrosPenyuntingCécile DecugisDistributorCompagnie Française de Distribution CinématographiqueTanggal rilis 15 Mei 1969 (1969-05-15) Durasi110 menitNegaraPrancisBahasaPrancis My Night at Maud's (bahasa Prancis: Ma nuit chez Maud) adalah sebuah film drama Prancis 19...

 

 

Якутский походОсновной конфликт: Гражданская война в России Якутские восстания Дата сентябрь 1921 — 19 июня 1923 Место Якутская область, ЯАССР, Приморская область, Охотское море Причина Недовольство местного населения политикой большевиков, проведение трудовой мобилизаци�...

 

 

Strategic operation conducted during Operation Barbarossa Battle of Białystok–MinskPart of Operation Barbarossa during the Eastern Front of World War IIRuins of Minsk, July 1941Date22 June – 9 July 1941LocationByelorussian Soviet Socialist Republic, Soviet UnionResult German victoryBelligerents  Germany  Soviet UnionCommanders and leaders Fedor von Bock Hermann Hoth Günther von Kluge Heinz Guderian Adolf Strauss Maximilian von Weichs Dmitry Pavlov  Vladimir Klimovskikh...

This biography of a living person needs additional citations for verification. Please help by adding reliable sources. Contentious material about living persons that is unsourced or poorly sourced must be removed immediately from the article and its talk page, especially if potentially libelous.Find sources: Pieter Kuijpers – news · newspapers · books · scholar · JSTOR (June 2018) (Learn how and when to remove this message) Pieter KuijpersBornPieter Ku...

 

 

Canadian radio personality and musician (born 1968) Nardwuar the Human ServietteNardwuar at Rolling Loud in 2019Background informationBirth nameJohn RuskinAlso known asNardwuar[1]Born (1968-07-05) July 5, 1968 (age 56)Vancouver, British Columbia, CanadaGenresGarage rockpunk rockOccupationsJournalistsingerkeyboardistInstrumentsVocalskeyboardsYears active1986–presentLabelsNardwuarAlternative TentaclesMintMember ofThe EvaporatorsWebsitenardwuar.comMusical artist Nardwuar the Human...