Toxochelys

Toxochelys Temporal range: Late Cretaceous

Fossil specimen, Houston Museum of Natural Science
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Suborder:	Cryptodira
Clade:	Panchelonioidea
Family:	†Toxochelyidae Baur, 1895
Genus:	†Toxochelys Cope, 1873
Species
T. latiremis Cope, 1873 (type) T. moorevillensis Zangerl, 1953

Toxochelys (/toʊxʌkɛliːz/) is an extinct genus of marine turtle from the Late Cretaceous period. It is the most commonly found fossilized turtle species in the Smoky Hill Chalk, in western Kansas.^[1]

Description

Toxochelys had carapace about 90 centimetres (3.0 ft) in length.^[2] Two species in the genus are recognized, Toxochelys latiremis and Toxochelys moorevillensis.^[3] Phylogenetic analysis shows that Toxochelys belong to an extinct lineage of turtles transitional between modern sea turtles and other turtles.^[4]

Toxochelys bauri Williston, 1905, based on the skeleton YPM 1786, is a synonym of Ctenochelys stenoporus.^[5]

Phylogeny

Toxochelys latiremis in a cladogram from the analysis of Gentry et al., 2019:^[6]

Pan-Chelonioidea

†Angolachelonia

†Protostegidae

†Toxochelys latiremis

	Chelonioidea

	†Ctenochelyidae

References

^ www.oceansofkansas.com Retrieved on May 12, 2008
^ Hirayama, Ren (1994). "Phylogenetic systematics of chelonioid sea turtles". Island Arc. 3 (4): 270–284. doi:10.1111/j.1440-1738.1994.tb00116.x. ISSN 1038-4871.
^ Nicholls, E.L. 1988. New material of Toxochelys latiremis Cope, and a revision of the genus Toxochelys (Testudines, Chelonoidea). Journal of Vertebrate Paleontology, 8(2):181–187.
^ Kear BP, Lee MS (March 2006). "A primitive protostegid from Australia and early sea turtle evolution". Biol. Lett. 2 (1): 116–9. doi:10.1098/rsbl.2005.0406. PMC 1617175. PMID 17148342.
^ R. Zangerl. 1953. The vertebrate fauna of the Selma Formation of Alabama. Part IV. The turtles of the family Toxochelyidae. Fieldiana: Geology Memoirs 3(4):145–277.
^ Andrew D. Gentry; Jun A. Ebersole; Caitlin R. Kiernan (2019). "Asmodochelys parhami, a new fossil marine turtle from the Campanian Demopolis Chalk and the stratigraphic congruence of competing marine turtle phylogenies". Royal Society Open Science. 6 (12): Article ID 191950. Bibcode:2019RSOS....691950G. doi:10.1098/rsos.191950. PMC 6936288. PMID 31903219.

Further reading

Cope, E. D. 1873. [On Toxochelys latiremis]. Proceedings of the Academy of Natural Sciences of Philadelphia 25:10.
Hay, O. P. 1896. On the skeleton of Toxochelys latiremis. Publ. Field Columbian Museum, Zoological Ser. (later Fieldiana: Zoology), 1(5):101–106, pls. 14 &15.
Case, E.C. 1898. Toxochelys. The University Geological Survey of Kansas, Part IV. 4:370–385. pls. 79–84.
Hay, O.P. 1905. A revision of the species of the family of fossil turtles called Toxochelyidae, with descriptions of two new species of Toxochelys and a new species of Porthochelys. Bulletin of the American Museum of Natural History 21(10):177–185.
Druckenmiller, P. S., A. J. Daun, J. L. Skulan and J. C. Pladziewicz. 1993. Stomach contents in the upper Cretaceous shark Squalicorax falcatus. Journal of Vertebrate Paleontology. 13(supplement. to no. 3):33A.
Carrino, M.H. 2007. Taxonomic comparison and stratigraphic distribution of Toxochelys (Testudines: Cheloniidae) of South Dakota. pp. 111–132 in Martin, J.E. and Parris D.C. (eds.), The Geology and Paleontology of the Late Cretaceous Marine Deposits of the Dakotas. Geological Society of America, Special Paper 427.
Konuki, R. 2008. Biostratigraphy of sea turtles and possible bite marks on a Toxochelys (Testudine, Chelonioidea) from the Niobrara Formation (Late Santonian), Logan County, Kansas and paleoecological implications for predator–prey relationships among large marine vertebrates. Unpublished Masters thesis, Fort Hays State University, Hays, Kansas, 141 pp, Appen. I-VI.
Matzke, A.T. 2008. A juvenile Toxochelys latiremis (Testudines, Cheloniidae) from the Upper Cretaceous Niobrara Formation of Kansas, USA. Neues Jahrbuch für Geologie und Paläontologie. Abhandlungen 249(3):371–380.
Matzke, A.T. 2009. Osteology of the skull of Toxochelys (Testudines, Chelonioidea) (with 25 text-figures). Palaeontographica Abteilung A 288(4):93–150.

External links

Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Diapsida
Order: Testudines

Genus

Chelonioidea
(Sea turtles)

Cheloniidae	†Allopleuron Caretta †Carolinochelys Chelonia †Eochelone Eretmochelys †Gigantatypus †Glarichelys †Itilochelys Lepidochelys †Mexichelys †Miocaretta Natator †Pacifichelys †Syllomus †Tasbacka
Dermochelyidae	†Arabemys †Corsochelys †Cosmochelys Dermochelys †Eosphargis †Mesodermochelys †Psephophorus
	†Euclastes †Peritresius †Procolpochelys †Protosphargis †Puppigerus

Dermatemydidae	Dermatemys
Kinosternidae	Claudius †Hoplochelys Kinosternon Staurotypus Sternotherus

Emydidae	†Acherontemys Chrysemys Clemmys Deirochelys Emys Actinemys Emydoidea Glyptemys Graptemys Malaclemys Pseudemys Terrapene Trachemys †Wilburemys
Geoemydidae	Batagur †Banhxeochelys Cuora Cyclemys Geoclemys Geoemyda Hardella Heosemys Leucocephalon Malayemys Mauremys Melanochelys Morenia Notochelys Orlitia Pangshura Rhinoclemmys Sacalia Siebenrockiella Vijayachelys
Platysternidae	Platysternon
Testudinidae	Aldabrachelys Astrochelys Centrochelys Chelonoidis Chersina Cylindraspis †Cymatholcus †Floridemys Geochelone Gopherus †Hadrianus †Hesperotestudo Homopus Indotestudo Kinixys Malacochersus Manouria †Megalochelys †Oligopherus Psammobates Pyxis †Solitudo Stigmochelys †Stylemys Testudo

Carettochelyidae	†Allaeochelys †Anosteira Carettochelys
Trionychidae	Amyda Apalone †Axestemys Chitra Cyclanorbis Cycloderma †Drazinderetes Dogania †Gilmoremys †Hutchemys †Khunnuchelys Lissemys Nilssonia Palea †Palaeoamyda Pelochelys Pelodiscus Rafetus Trionyx
	†Basilochelys †Sinaspideretes

Chelydridae	Chelydra †Chelydrops †Chelydropsis †Emarginachelys †Macrocephalochelys Macrochelys †Planiplastron †Protochelydra
†Nanhsiungchelyidae	†Anomalochelys †Basilemys †Jiangxichelys
†Protostegidae	†Alienochelys †Archelon †Atlantochelys †Bouliachelys †Calcarichelys †Cratochelone †Desmatochelys †Iserosaurus †Notochelone †Ocepechelon †Pneumatoarthrus †Protostega †Rhinochelys †Santanachelys †Terlinguachelys
	†Adocus †Argillochelys †Bashuchelys †Ctenochelys †Prionochelys †Toxochelys

†Araripemydidae	†Araripemys
†Bothremydidae	†Araiochelys †Arenila †Azabbaremys †Bothremys †Cearachelys †Chedighaii †Chupacabrachelys †Eotaphrosphys †Foxemys †Galianemys †Ilatardia †Inaechelys †Itapecuruemys †Jainemys †Kinkonychelys †Kurmademys †Labrostochelys †Nigeremys †Phosphatochelys †Polysternon †Puentemys †Rosasia †Rhothonemys †Sankuchemys †Taphrosphys †Ummulisani †Zolhafah
Chelidae	Acanthochelys Chelodina Chelus Elseya Elusor Emydura Hydromedusa †Lomalatachelys Mesoclemmys Myuchelys Phrynops Platemys †Prochelidella Pseudemydura Rheodytes Rhinemys †Yaminuechelys
Pelomedusidae	Pelomedusa Pelusios
Podocnemididae	†Albertwoodemys †Bauruemys †Brontochelys †Caninemys †Carbonemys †Cerrejonemys †Cordichelys Erymnochelys †Lapparentemys †Latentemys Peltocephalus Podocnemis †Stupendemys
†Sahonachelyidae	†Sahonachelys †Sokatra

	†Caribemys †Caririemys †Tacuarembemys

Phylogenetic arrangement of turtles based on Turtles of the World 2017 Update: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status. † = extinct.

See also List of Testudines families

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