Howellia

Howellia

Vulnerable  (NatureServe)[1]

Delisted (ESA)[2]
Scientific classification Edit this classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Asterids
Order: Asterales
Family: Campanulaceae
Subfamily: Lobelioideae
Genus: Howellia
A.Gray
Species:
H. aquatilis
Binomial name
Howellia aquatilis
A.Gray

Howellia aquatilis, the water howellia, is a small plant in the family Campanulaceae that was considered a federally threatened species between 1994[3] and 2021 under the Endangered Species Act of 1973.[2][4] It is the only species in the monotypic genus Howellia.[5] Thomas Jefferson Howell and Joseph Howell discovered the species in 1878.[6]

Distribution

The water howellia is found in large area of the Pacific Northwest region of the United States. It can be found in California, Idaho, Montana, and Washington, and it is known historically from Oregon.[7]

The plant grows in wetland habitat surrounded by forests, which provide organic material.[5] Associated tree species include Populus trichocarpa (California poplar), P. tremuloides (quaking aspen), and Fraxinus latifolia (Oregon ash).[5] The water bodies are located in glacial potholes, river oxbows, ephemeral ponds, flood plains and other areas that fill with water periodically as snow melts and spring rain falls.[5]

Biology

The plant produces two types of flowers. Submerged cleistogamous flowers remain closed and self-pollinate, and flowers that bloom above the surface of the water open into white blossoms and may cross with other individuals.[5] Because it often pollinates itself, the species has a low genetic diversity.[5] Seeds are produced in the water but they require open air for germination, so they sprout when the wetland has dried in the fall.[5]

Conservation

Threats to this species include loss of habitat to logging and draining and conversion to urban use, alteration of the local hydrology by placement of dams and dikes, deposition of silt, livestock activity, introduced species of plants, low genetic diversity, and climate change.[8]

References

  1. ^ NatureServe (3 March 2023). "Howellia aquatilis". NatureServe Network Biodiversity Location Data accessed through NatureServe Explorer. Arlington, Virginia: NatureServe. Retrieved 3 April 2023.
  2. ^ a b 86 FR 31955
  3. ^ 59 FR 35860
  4. ^ "Endangered and Threatened Wildlife and Plants; Removal of Howellia aquatilis (Water Howellia) From the List of Endangered and Threatened Plants". Federal Register. 2019-10-07. Retrieved 2019-10-06.
  5. ^ a b c d e f g "Center for Plant Conservation". Archived from the original on 2011-08-07. Retrieved 2011-04-16.
  6. ^ Engeman, Richard H. (2009). The Oregon Companion: An Historical Gazetteer of The Useful, The Curious, and The Arcane. Portland, Oregon: Timber Press. p. 184. ISBN 978-0-88192-899-0.
  7. ^ USFWS Species Profile.
  8. ^ USFS Wildflowers.

Read other articles:

Pakistan Navy

Maritime service branch of the Pakistan Armed Forces Pakistan Navyپاکستان بحریہCrest of Pakistan NavyFounded14 August 1947 (1947-08-14)[Note 1](76 years, 6 months ago)Country PakistanTypeNavyRole Naval warfare Maritime security Coastal defence Second-strike capability[1] Size[2]: 73 [3]: 33 [4][5][6] 50,000 active-duty personnel 50,000 reserve force 18,000 Marines ...

 

 

Aiden Ashley

Artikel ini sebatang kara, artinya tidak ada artikel lain yang memiliki pranala balik ke halaman ini.Bantulah menambah pranala ke artikel ini dari artikel yang berhubungan atau coba peralatan pencari pranala.Tag ini diberikan pada Desember 2023. Aiden AshleyLahirAiden Ashley28 Februari 1990 (umur 34)Arizona, Amerika SerikatKebangsaanAmerikaTinggi5 ft 3 in (1,60 m)Berat121 pon (55 kg) Aiden Ashley (lahir 28 Februari 1990) adalah seorang pemeran pornografi asal Ame...

 

 

Daftar runner-up Kejuaraan Dunia Pembalap Formula Satu

Artikel ini sebatang kara, artinya tidak ada artikel lain yang memiliki pranala balik ke halaman ini.Bantulah menambah pranala ke artikel ini dari artikel yang berhubungan atau coba peralatan pencari pranala.Tag ini diberikan pada Desember 2023. Formula Satu Musim berjalan Formula Satu musim 2023 Artikel terkait Sejarah Formula Satu Balapan Formula Satu Regulasi Formula Satu Mobil Formula Satu Mesin Formula Satu Ban Formula Satu Daftar Pembalap (Pemenang GPPembalap polePembalap lap tercepat J...

Heroes (We Could Be)

Heroes (We Could Be)Singel oleh Alesso featuring Tove Lodari album ForeverDirilis22 Agustus 2014Direkam2012–2014GenreProgressive houseDurasi3:30 (Radio Edit)5:39 (Extended Mix)LabelDef JamPenciptaAlessandro LindbladEbba NilssonDavid JonesBrian EnoProduserAlessoKronologi singel Alesso Tear the Roof Up (2014) Heroes (We Could Be) (2014) Cool (2015) Kronologi singel Tove Lo Stay High(2014) Heroes (We Could Be)(2014) Not on Drugs(2014) Video musikHeroes di YouTube Heroes (We Could Be)...

 

 

Keuskupan Agung Chambéry–Saint-Jean-de-Maurienne–Tarentaise

Keuskupan Agung Chambéry, Saint-Jean-de-Maurienne, dan TarentaiseArchidioecesis Camberiensis, Maruianensis et TarantasiensisArchidiocèse de Chambéry, Saint-Jean-de-Maurienne et TarentaiseKatolik Katedral ChambéryLokasiNegaraPrancisProvinsi gerejawiLyonStatistikLuas7.460 km2 (2.880 sq mi)Populasi- Total- Katolik(per 2013)432.000397,700 (92.1%)InformasiDenominasiKatolik RomaGereja sui iurisGereja LatinRitusRitus RomaPendirianDisatukan: 26 April 1966Katedra...

 

 

Hamad bin Khalifa al-Tsani

Hamad bin Khalifa Al-Thaniحمد بن خليفة آل ثانيEmir QatarBerkuasa27 June 1995 – 25 Juni 2013PendahuluKhalifa bin Hamad al-ThaniPenggantiTamim bin Hamad al-ThaniPerdana Menteri Lihat daftar Abdullah bin Khalifa al-ThaniHamad bin Jassim bin Jaber al-Thani Perdana Menteri QatarBerkuasa27 Juni 1995 – 29 Oktober 1996PendahuluKhalifa bin Hamad al-ThaniPenerusAbdullah bin Khalifa al-ThaniInformasi pribadiKelahiran1952Doha, QatarWangsaAl-TsaniAyahKhalifa bin Hamad al-TsaniIbuSheikh...

Official residence

Residence of head of state An official residence is a residence designated by an authority and assigned to an official (such as a head of state, head of government, governor, or other senior figures), and may not always be the same place where the office holder conducts their official functions or lives.[1][2][3][4] List of official residences, by country Afghanistan Arg (Cabinet) Albania Prime Minister's Office Pallati i Brigadave Ish-Blloku (former residence ...

 

 

Sha (Mongolic)

Letter of related and vertically oriented alphabets used to write Mongolic and Tungusic languages Sha is a letter of related and vertically oriented alphabets used to write Mongolic and Tungusic languages.[1]: 549–551  Mongolian language Look up ᠱ in Wiktionary, the free dictionary. Main articles: Mongolian script, Mongolian writing systems, and Mongolian language ShaThe Mongolian scriptMongolian vowelsᠠ​aᠡ​eᠢ​iᠣ​oᠤ​uᠥ̴...

 

 

Slovakia men's national basketball team

This article is about the men's team. For the women's team, see Slovakia women's national basketball team. SlovakiaFIBA ranking60 (1 March 2024)[1]Joined FIBA1993FIBA zoneFIBA EuropeNational federationSBACoachAramis NaglićNickname(s)Repre(The Representatives)FIBA World CupAppearancesNoneEuroBasketAppearancesNone Home Away First international Bosnia 97–69 Slovakia (Wrocław, Poland; 30 May 1993)Biggest win Slovakia 100–39 Malta (Levice, Slovakia; 7 September 20...

Government House, Suva

Fiji State HouseThe State House, viewed from Queen Elizabeth DriveFormer namesGovernment HouseAlternative namesPresidential PalaceGeneral informationTypeOfficial residenceArchitectural styleGeorgianAddressQueen Elizabeth DriveTown or citySuvaCountryFijiCoordinates18°09′07″S 178°25′34″E / 18.151892°S 178.426140°E / -18.151892; 178.426140Current tenantsWiliame Katonivere, President of FijiCompleted1928ClientColony of FijiOwnerGovernment of Fiji The State Hous...

 

 

Meikyū Love Song

Meikyū Love Song(迷宮ラブソング)Lagu oleh Arashidari album PopcornDirilis02 November 2011 (2011-11-02)FormatCD, CD + DVDDirekam2011GenrePopLabelJ StormKronologi singel Lotus (2011) Meikyū Love Song(迷宮ラブソング) (2011) Wild at Heart (2012) Meikyū Love Song (迷宮ラブソング) adalah sebuah lagu yang dinyanyikan oleh boyband Arashi. Lagu ini dirilis pada tanggal 2 November 2011 oleh label rekaman mereka J Storm. Meikyū Love Song digunakan untuk lagu tema drama Nazo...

 

 

The Michael J. Fox Show

American television sitcom (2013–2014) The Michael J. Fox ShowGenreSitcomCreated by Will Gluck Sam Laybourne Starring Michael J. Fox Betsy Brandt Wendell Pierce Katie Finneran Juliette Goglia Conor Romero Jack Gore Ana Nogueira ComposerJeff CardoniCountry of originUnited StatesOriginal languageEnglishNo. of seasons1No. of episodes22 (7 unaired in the U.S.)ProductionExecutive producers Alex Reid Richard Schwartz Sam Laybourne Michael J. Fox Will Gluck Camera setupSingle-cameraRunning time20 ...

Мелодичный дэт-метал

Мелодичный дэт-метал(мелодэт) Направление метал и экстремальный метал Истоки хеви-метал, трэш-метал, дэт-метал[1], пауэр-метал Время и место возникновения Начало – середина 1990-х,  Англия,  Швеция Музыкальные инструменты Вокал (гроулинг, харш, скриминг, чистый вока�...

 

 

Pembicaraan Pengguna:Raksasabonga

Halo, Raksasabonga. Selamat datang di Wikipedia bahasa Indonesia! Memulai Bacalah halaman Pengantar Wikipedia terlebih dahulu. Baca juga informasi tentang berkontribusi di Wikipedia. Tuliskan juga sedikit profil Anda di Pengguna:Raksasabonga, halaman pribadi Anda, agar kami dapat lebih mengenal Anda. Lihat pula aturan yang disederhanakan sebelum melanjutkan. Tips Selalu tanda tangani pertanyaan Anda di Warung Kopi atau halaman pembicaraan dengan mengetikkan ~~~~ pada akhir kalimat Anda. Jang...

 

 

مساهمون لعلم الكون

يفتقر محتوى هذه المقالة إلى الاستشهاد بمصادر. فضلاً، ساهم في تطوير هذه المقالة من خلال إضافة مصادر موثوق بها. أي معلومات غير موثقة يمكن التشكيك بها وإزالتها. (فبراير 2019) هذه المقالة بها مشكلات متعدِّدة. فضلًا ساعد في تحسينها أو ناقش هذه المشكلات في صفحة النقاش. لم تُحدد المُ�...

الجزائر في الألعاب الأولمبية

يفتقر محتوى هذه المقالة إلى الاستشهاد بمصادر. فضلاً، ساهم في تطوير هذه المقالة من خلال إضافة مصادر موثوق بها. أي معلومات غير موثقة يمكن التشكيك بها وإزالتها. (فبراير 2016) الجزائر في الألعاب الأولمبية علم الجزائر رمز ل.أ.د.  ALG ل.أ.و. اللجنة الأولمبية الجزائرية تاريخ �...

 

 

卡塔尔首相

卡塔尔首相卡塔尔国徽現任穆罕默德·本·阿卜杜勒拉赫曼·阿勒薩尼自2023年3月7日在任官邸Green Palace任命者卡塔尔埃米尔首任哈利法·本·哈马德阿勒萨尼设立1970年5月29日 卡塔尔首相 (阿拉伯语：رئیس الوزرا القطری‎是卡塔尔的政府首脑，是卡塔尔国内仅次于埃米尔的第二位最有权力的人物。 1970年5月29日，第十任埃米尔哈利法·本·哈马德·阿勒萨尼创建了首相一�...

 

 

Computational neuroscience

Branch of neuroscience This article may be too technical for most readers to understand. Please help improve it to make it understandable to non-experts, without removing the technical details. (March 2014) (Learn how and when to remove this message) Computational neuroscience (also known as theoretical neuroscience or mathematical neuroscience) is a branch of neuroscience which employs mathematics, computer science, theoretical analysis and abstractions of the brain to understand t...

Chevrolet Greenbrier

Si ce bandeau n'est plus pertinent, retirez-le. Cliquez ici pour en savoir plus. Cette section ou cet article est une traduction incomplète (juillet 2021). Vous pouvez modifier la page pour effectuer la traduction. Si ce bandeau n'est plus pertinent, retirez-le. Cliquez ici pour en savoir plus. Certaines informations figurant dans cet article ou cette section devraient être mieux reliées aux sources mentionnées dans les sections « Bibliographie », « Sources » ou «...

 

 

National League 2019-2020 (Gibilterra)

National League 2019-2020 Competizione National League Sport Calcio Edizione 121ª Organizzatore GFA Date dal 12 agosto 2019al 29 febbraio 2020 Luogo  Gibilterra Partecipanti 12 Risultati Vincitore non assegnato Retrocessioni nessuna Statistiche Miglior marcatore Juanfri (24) Incontri disputati 104 Gol segnati 481 (4,63 per incontro) Cronologia della competizione 2020-2021 Manuale La National League 2019-2020 è stata la 1ª edizione dell'omonima competizione a seguito...