Cladistia
Clade of ray-finned fishes
Cladistia is a clade of bony fishes whose only living members are the bichirs of tropical Africa.[ 1] Their major synapomorphies are a heterocercal tail in which the dorsal fin has independent rays, and a posteriorly elongated parasphenoid .
Cladistia are the earliest diverging branch of living Actinopterygii , and are thought to have diverged from the Actinopteri , the group which includes all other living ray finned fish, by the Carboniferous .[ 2] However, the fossil range for the only extant order (Polypteriformes ) is comparatively young, only reaching as far back as the mid-Cretaceous of South America and Africa, and the two extant genera of bichir only diverged around the Miocene .[ 3]
Aside from bichirs, other extinct fish groups thought to be members of the group include the Scanilepiformes , known from Triassic (and possibly Permian [ 4] ) of the Northern Hemisphere.[ 5] [ 2] [ 6] The Guildayichthyiformes of Carboniferous North America are also sometimes considered cladistians, but this is thought to be dubious, with other authorities placing them as a stem-group to Neopterygii .[ 6] [ 7]
Taxonomy
Based on work done by Near et al (2017) and Lund (2000):[ 8]
Order Polypteriformes Bleeker 1859
Genus †Latinopollia Meunier and Gayet 1998
Genus †Dagetella Gayet & Meunier 1991
Genus †Bartschichthys Gayet & Meunier 1996 [Bartschia Gayet & Meunier 1996 non Rehder 1943 ]
Genus †Sudania Werner & Gayet 1997
Genus †Saharichthys Werner & Gayet 1997
Genus †Sainthilairia Gayet & Meunier 1996
Species †S. beccussiformis Gayet & Meunier 1996
Species †S. elongata Werner & Gayet 1997
Species †S. falciformis Gayet & Meunier 1996
Species †S. grandis Gayet & Meunier 1996
Species †S. intermedia Werner & Gayet 1997
Genus †Inbecetemia Werner & Gayet 1997
Genus †Nagaia Werner & Gayet 1997
Family Polypteridae Lacépède 1803 sensu stricto
Genus †Bawitius Grandstaff et al. 2012
Genus †Serenoichthys Dutheil 1999
Genus Erpetoichthys Smith 1865 [Calamoichthys Smith 1866 ]
Genus Polypterus Lacepède 1803
Species Polypterus ansorgii Boulenger , 1910
Species Polypterus bichir Lacépède , 1803
Species Polypterus congicus Boulenger , 1898
Species Polypterus delhezi Boulenger , 1899
Species Polypterus endlicherii Heckel , 1847
Species Polypterus mokelembembe Schliewen & Schäfer , 2006
Species Polypterus ornatipinnis Boulenger , 1902
Species Polypterus palmas Ayres , 1850
Species Polypterus polli J. P. Gosse , 1988
Species Polypterus retropinnis Vaillant , 1899
Species Polypterus senegalus Cuvier , 1829
Species Polypterus teugelsi Britz , 2004
Species Polypterus weeksii Boulenger , 1898
Order †Scanilepiformes Sytchevskaya, 1999 [ 2] [ 9] [ 10]
Disputed members
References
^ Lecointre G, Le Guyader H (2007). The Tree of Life: A Phylogenetic Classification . Harvard University Press Reference Library. ISBN 978-0-674-02183-9 .
^ a b c Giles, Sam; Xu, Guang-Hui; Near, Thomas J.; Friedman, Matt (2017). "Early members of 'living fossil' lineage imply later origin of modern ray-finned fishes" . Nature . 549 (7671): 265–268. doi :10.1038/nature23654 . ISSN 1476-4687 .
^ Near, Thomas J.; Dornburg, Alex; Tokita, Masayoshi; Suzuki, Dai; Brandley, Matthew C.; Friedman, Matt (2014-01-02). "BOOM AND BUST: ANCIENT AND RECENT DIVERSIFICATION IN BICHIRS (POLYPTERIDAE: ACTINOPTERYGII), A RELICTUAL LINEAGE OF RAY-FINNED FISHES" . Evolution . 68 (4): 1014–1026. doi :10.1111/evo.12323 . ISSN 0014-3820 .
^ Bakaev, A. S.; Kogan, I. (2022). "Squamation of the Permian actinopterygian Toyemia Minich, 1990: evenkiid (Scanilepiformes) affinities and implications for the origin of polypteroid scales" . www.geology.cz . Retrieved 2024-02-28 .
^ Argyriou T, Giles S, Friedman M, Romano C, Kogan I, Sánchez-Villagra MR (November 2018). "Internal cranial anatomy of Early Triassic species of †Saurichthys (Actinopterygii: †Saurichthyiformes): implications for the phylogenetic placement of †saurichthyiforms" . BMC Evolutionary Biology . 18 (1): 161. doi :10.1186/s12862-018-1264-4 . PMC 6211452 . PMID 30382811 .
^ a b Near, Thomas J; Thacker, Christine E (18 April 2024). "Phylogenetic classification of living and fossil ray-finned fishes (Actinopterygii)" . Bulletin of the Peabody Museum of Natural History . 65 . doi :10.3374/014.065.0101 .
^ "The new Actinopterygian order Guildayichthyiformes from the Lower Carboniferous of Montana (USA)" . Geodiversitas . 22 (2): 171–206. 2000.
^ Mikko Haaramo. "Cladistia – bichirs and relatives" . Mikko's Phylogeny Archive . Retrieved 2013-10-26 .
^ "PBDB" . paleobiodb.org . Retrieved 2024-02-28 .
^ Xu, Guang-Hui; Gao, Ke-Qin (2011). "A new scanilepiform from the Lower Triassic of northern Gansu Province, China, and phylogenetic relationships of non-teleostean Actinopterygii: EARLY TRIASSIC SCANILEPIFORM FROM CHINA" . Zoological Journal of the Linnean Society . 161 (3): 595–612. doi :10.1111/j.1096-3642.2010.00645.x .