Bolboceratidae is a family of beetle.[1] It was historically treated as a subfamily of the earth-boring dung beetles, or "dor beetles" (family Geotrupidae), but has been considered a separate family by many authors since 1995.[2] Some recent classifications have persisted in treating bolboceratids as a subfamily (e.g.[3][4]) but these classifications are contradicted by recent phylogenetic studies of relationships indicating that bolboceratids are not closely related to geotrupids (e.g., that bolboceratids are more closely related to Pleocomidae and Passalidae[5]).
^Smith, Andrew B. T. (2006). "A Review of the Family-group Names for the Superfamily Scarabaeoidea (Coleoptera) with Corrections to Nomenclature and a Current Classification". The Coleopterists Bulletin. 60 (sp5). Coleopterists Society: 144–204. doi:10.1649/0010-065x(2006)60[144:arotfn]2.0.co;2. ISSN0010-065X.
^Mckenna, Duane D.; Wild, Alexander L.; Kanda, Kojun; Bellamy, Charles L.; Beutel, Rolf G.; et al. (2015). "The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution". Systematic Entomology. 40 (4): 835–880. Bibcode:2015SysEn..40..835M. doi:10.1111/syen.12132. hdl:10057/11540. ISSN0307-6970.
