Paleontology or palaeontology is the study of prehistoriclife forms on Earth through the examination of plant and animal fossils.[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2009.
Three new species of extinct Octopoda discovered in 2009. The species – Keuppia hyperbolaris, Keuppia levante, and Styletoctopus annae – lived about 95 million years ago, and bear a strong resemblance to modern octopuses, suggesting that the Octopoda order has remained relatively unchanged for tens of millions of years. The fossils included evidence of arms, muscles, rows of suckers, ink, and internal gills. The discovery was made by a team led by Dirk Fuchs of the Freie University, which is located at Berlin, Germany.[2] The fossils were found at Hakel and Hadjoula, Lebanon.[3] Various new ammonoid taxa were named, including Ivoites.
A member of Percoidei. The type species is "Serranus" transsylvanicus Bohm (1942).
Shimada, K.; Everhart, M.J. (2009). "First record of Anomoeodus (Osteichthyes: Pycnodontiformes) from the Upper Cretaceous Niobrara Chalk of western Kansas". Transactions of the Kansas Academy of Science. 112 (1/2): 98–102. doi:10.1660/062.112.0212. S2CID83951062.
A basal member of Testudinata. The type species is C. tenertesta. The genus Chinlechelys was considered to be a junior synonym of the genus Proganochelys by Joyce (2017), though the author maintained C. tenertesta as a distinct species within the latter genus.[30]
A study is published that proposes that females from certain taxa use ornaments as a criterion for mate choice because other dimorphic structures, like biological "weaponry" could be used to coerce or force them to mate.[50]
A study concludes that biotic factors have more pronounced local and short term evolutionary impacts than abiotic factors, which in turn have a more pronounced effect through time and on biodiversity as a whole.[51]
Extinction
A study noting the effects of the KT mass extinction on Earth's modern biota is published.[52]
Geology
Zhang, H.; Wei, Z.-L.; Liu, X.-M.; Li, D. (2009). "Constraints on the age of the Tuchengzi Formation by LA-ICP-MS dating in northern Hebei-western Liaoning, China". Science China Earth Sciences. 52 (4): 461–470. Bibcode:2009ScChD..52..461Z. doi:10.1007/s11430-009-0052-9. S2CID129751106.
Ichnology
Bedatou, E.; Melchor, R.N.; Genise, J.F. (2009). "Complex palaeosol ichnofabrics from Late Jurassic-Early Cretaceous volcaniclastic successions of central Patagonia, Argentina". Sedimentary Geology. 218 (1–4): 74–102. Bibcode:2009SedG..218...74B. doi:10.1016/j.sedgeo.2009.04.005.
Paleobiogeography
Pereda-Suberbiola X (2009). "Biogeographical affinities of Late Cretaceous continental tetrapods of Europe: a review". Bulletin de la Société Géologique de France. 180 (1): 57–71. doi:10.2113/gssgfbull.180.1.57.
Paleoecology
Nicolas M., Rubidge B.S. (2009). "Changes in Permo-Triassic terrestrial tetrapod ecological representation in the Beaufort Group (Karoo Supergroup) of South Africa". Lethaia. 43: 45–59. doi:10.1111/j.1502-3931.2009.00171.x.
Footnotes
Complete author list
As science becomes more collaborative, papers with large numbers of authors are becoming more common. To prevent the deformation of the tables, these footnotes list the contributors to papers that erect new genera and have many authors.
^Gini-Newman, Garfield; Graham, Elizabeth (2001). Echoes from the past: world history to the 16th century. Toronto: McGraw-Hill Ryerson Ltd. ISBN9780070887398. OCLC46769716.
^De Baets, K.; Klug, C.; Korn, D. (2009). "Anetoceratinae (Ammonoidea, Early Devonian) from the Eifel and Harz Mountains (Germany), with a revision of their genera". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 252 (3): 361–376. doi:10.1127/0077-7749/2009/0252-0361.
^N.‐Z. Wang; X. Zhang; M. Zhu; W.‐J. Zhao (2009). "A new articulated hybodontoid from Late Permian of northwestern China". Acta Zoologica. 90 (s1): 159–170. doi:10.1111/j.1463-6395.2008.00382.x.
^ abEileen D. Grogan; Richard Lund (2009). "Two new iniopterygians (Chondrichthyes) from the Mississippian (Serpukhovian) Bear Gulch Limestone of Montana with evidence of a new form of chondrichthyan neurocranium". Acta Zoologica. 90 (s1): 134–151. doi:10.1111/j.1463-6395.2008.00371.x.
^Davit Vasilyan; Bettina Reichenbacher; Giorgio Carnevale (2009). "A new fossil Aphanius species from the Upper Miocene of Armenia (Eastern Paratethys)". Paläontologische Zeitschrift. 83 (4): 511–519. doi:10.1007/s12542-009-0034-4. S2CID129643131.
^Brian Choo; John A. Long; Katherine Trinajstic (2009). "A new genus and species of basal actinopterygian fish from the Upper Devonian Gogo Formation of Western Australia". Acta Zoologica. 90 (s1): 194–210. doi:10.1111/j.1463-6395.2008.00370.x.
^Snitting, D. (2009). "Heddleichthys- a new tristichopterid genus from the Dura Den Formation, Midland Valley, Scotland (Famennian, Late Devonian)". Acta Zoologica. 90: 273–284. doi:10.1111/j.1463-6395.2008.00376.x.
^Louis Taverne (2009). "Ridewoodichthys, a new genus for Brychaetus caheni from the marine Paleocene of Cabinda (Africa): re-description and comments on its relationships within the Osteoglossidae (Teleostei, Osteoglossomorpha)". Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre. 79: 147–153.
^ abBáez, Ana M.; Moura, Geraldo J.B.; Gómez, Raúl O.; Ana M. Báez; Geraldo J.B. Moura; Raúl O. Gómez (2009). "Anurans from the Lower Cretaceous Crato Formation of northeastern Brazil: implications for the early divergence of neobatrachians". Cretaceous Research. 30 (4): 829–846. doi:10.1016/j.cretres.2009.01.002.
^Anderson, J.S.; Scott, D.; Reisz, R.R. (2009). "Nannaroter mckinziei, a new ostodolepid 'microsaur' (Tetrapoda, Lepospondyli, Recumbirostra) from the Early Permian of Richards Spur (Ft. Sill), Oklahoma". Journal of Vertebrate Paleontology. 29 (2): 379–388. doi:10.1671/039.029.0222. S2CID130420068.
^Skutschas, Pavel P.; Pavel P. Skutschas (2009). "Re-Evaluation of Mynbulakia Nesov, 1981 (Lissamphibia: Caudata) and Description of a New Salamander Genus from the Late Cretaceous of Uzbekistan". Journal of Vertebrate Paleontology. 29 (3): 659–664. doi:10.1671/039.029.0326. S2CID85024012.
^Zhang, Guilin; Wang, Yuan; Jones, Marc E.H.; Evans, Susan E.; Guilin Zhang; Yuan Wang; Marc E.H. Jones; Susan E. Evans (2009). "A new Early Cretaceous salamander (Regalerpeton weichangensis gen. et sp. nov.) from the Huajiying Formation of northeastern China". Cretaceous Research. 30 (3): 551–558. doi:10.1016/j.cretres.2008.10.004.
^Klembara, Jozef (2009). "The skeletal anatomy and relationships of a new discosauriscid seymouriamorph from the lower Permian of Moravia (Czech Republic)". Annals of Carnegie Museum. 77 (4): 451–483. doi:10.2992/0097-4463-77.4.451. S2CID86629854.
^Vandermark, D.; Tarduno, J.A.; Brinkman, D.B.; Cottrell, R.D.; Mason, S. (2009). "New Late Cretaceous macrobaenid turtle with Asian affinities from the High Canadian Arctic: dispersal via ice-free polar routes". Geology. 37 (2): 183–186. Bibcode:2009Geo....37..183V. doi:10.1130/G25415A.1.
^Tong, H.; Claude, J.; Naksri, W.; Suteethorn, V.; Buffetaut, E.; Khansubha, S.; Wongko, K. & Yuangdetkla, P. (2009). "Basilochelys macrobios n. gen. and n. sp., a large cryptodiran turtle from the Phu Kradung Formation (latest Jurassic-earliest Cretaceous) of the Khorat Plateau, NE Thailand". In: Buffetaut, E.; Cuny, G.; Le Loeuff, J. & Suteethorn, V. (eds.). Late Palaeozoic and Mesozoic Ecosystems in SE Asia. Geological Society, London, Special Publications 315: 229-243.
^ abLyson, T.R.; Joyce, W.G. (2009). "A revision of Plesiobaena (Testudinoes: Baenidae) and an assessment of Baenid ecology across the K/T boundary". Journal of Paleontology. 83 (6): 833–853. doi:10.1666/09-035.1. S2CID85964417.
^Richard Franz; Shelley E. Franz (2009). "A new fossil land tortoise in the genus Chelonoidis (Testudines: Testudinidae) from the northern Bahamas, with an osteological assessment of other Neotropical tortoises". Bulletin of the Florida Museum of Natural History. 49 (1): 1–44.
^ abWalter G. Joyce; Tyler R. Lyson (2011). "New Material of Gilmoremys lancensis nov. comb. (Testudines: Trionychidae) from the Hell Creek Formation and the Diagnosis of Plastomenid Turtles". Journal of Paleontology. 85 (3): 442–459. doi:10.1666/10-127.1. S2CID129918006.
^Lyson, T.R.; Joyce, W.G. (2009). "A New Species of Palatobaena (Testudines: Baenidae) and a Maximum Parsimony and Bayesian Phylogenetic Analysis of Baenidae". Journal of Paleontology. 83 (3): 457–470. doi:10.1666/08-172.1. S2CID86045244.
^Druckenmiller, P.S.; Russell, A.P. (2009). "The new plesiosaurian genus Nichollssaura from Alberta, Canada: replacement name for the preoccupied genus Nichollsia". Journal of Vertebrate Paleontology. 29 (1): 276. doi:10.1080/02724634.2009.10010379. S2CID83847722.
^Hani Faig Kaddumi (2009). "On the remains of the first plesiosaur (Reptilia: Sauropterygia) from Harrana with a description of a new genus and species of Polycotylidae". In Hani Faig Kaddumi (ed.). Fossils of the Harrana Fauna and the Adjacent Areas. pp. 158–162.
^Head, Jason J.; Jonathan I. Bloch; Alexander K. Hastings; Jason R. Bourque; Edwin A. Cadena; Fabiany A. Herrera; P. David Polly; Carlos A. Jaramillo (2009). "Giant boid snake from the paleocene neotropics reveals hotter past equatorial temperatures". Nature. 457 (7230): 715–718. Bibcode:2009Natur.457..715H. doi:10.1038/nature07671. PMID19194448. S2CID4381423.