2004 in paleontology

List of years in paleontology (table)
In science
2001
2002
2003
2004
2005
2006
2007
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Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2004.

Protozoa

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Paleoleishmania[2]

Gen et sp nov

Valid

Poinar & Poinar

Albian

Burmese amber

 Myanmar

Oldest record of the protozoan family Trypanosomatidae,
Type species P. proterus

Paleoleishmania proterus

Fungi

newly named

Name Novelty Status Authors Age Unit Location Notes Images

Appianoporites[3]

Gen et sp nov

Valid

Smith, Currah, & Stockey

Eocene

Appian Way Flora.

 Canada

Extinct bracket fungus

Quatsinoporites[3]

Gen et sp nov

Valid

Smith, Currah, & Stockey

Barremian (Cretaceous)

Longarm Formation.

 Canada

Extinct bracket fungus.

Plants

Newly named plants

Name Novelty Status Authors Age Unit Location Notes Images

Fagus langevinii[4]

Sp nov

Valid

Manchester & Dillhoff

Ypresian

Tranquille Formation

 Canada
 British Columbia

A beech tree species

Fagus langevinii

Liquidambar changii[5]

Sp nov.

Valid

Pigg, Ickert-Bond, & Wen

Middle Miocene

"Yakima Canyon Flora"

 USA

A species of sweetgum

Liquidambar changii

Neviusia dunthornei[6]

Sp nov

Valid

DeVore, Moore, Pigg, & Wehr

Eocene
Ypresian

Okanagan Highlands
Allenby Formation

 Canada
 British Columbia

oldest and only extinct species of Neviusia

Palaeocarpinus dakotensis[7]

Sp nov

Valid

Manchester & Chen

Paleocene
Tiffanian

Fort Union Group
Sentinel Butte Formation

 USA
 North Dakota

A betulaceous fruit

Arthropoda

newly named arachnids

Name Novelty Status Authors Age Unit Location Notes Images

?Anyphops cortex[8]

sp nov

jr synonym[9]

Wunderlich

Unknown

Madagascar copal

 Madagascar

jr Synonym of Garcorops jadis

Garcorops jadis[10]

sp nov

Valid

Bosselaers

Unknown

Madagascar copal

 Madagascar

Possibly extant, but copal age is not determined

Newly named insects

Name Novelty Status Authors Age Unit Location Notes Images

Afromyrma[11]

Gen et sp nov

valid

Dlussky, Brothers & Rasnitsyn

Turonian

 Botswana

A Myrmicin ant, type species A. petrosa

Afropone[11]

Gen et sp nov

valid

Dlussky, Brothers & Rasnitsyn

Turonian

 Botswana

A ponerin ant, two species

Albicoccus[12]

Fam, gen et sp nov

valid

Koteja

Albian - Cenomanian

Burmese amber

 Myanmar

A scale insect, monotypic with Albicoccidae & A. dimai

Burmacoccus[12]

Fam, gen et sp nov

valid

Koteja

Albian - Cenomanian

Burmese amber

 Myanmar

A scale insect, monotypic with Burmacoccidae & B. danyi

Carebara antiqua[13]

Comb nov

valid

(Mayr, 1868)

Middle Eocene

Baltic amber

 Europe

Fossil myrmicine ant

Carebara antiqua

Electrostephanus janzeni[14]

Sp nov

jr synonym

Engel

Lutetian

Baltic Amber

 Russia

A stephanid wasp

Gerontoformica[15]

Gen et sp nov

valid

Nel & Perrault

Albian - Cenomanian

Charentese amber

 France

A stem group ant,
type species G. cretacica

Gerontoformica cretacica

Marmyan[12]

gen et sp nov

valid

Koteja

Cenomanian

Burmese amber

 Myanmar

A scale insect, monotypic M. barbarae

Microberotha[16]

Gen et sp nov

Valid

Archibald

Ypresian

Hat Creek Amber

 Canada

A Berothid lacewing

Palaeomyia[17]

Gen et sp nov

Valid

Poinar

Cenomanian

Burmese amber

 Myanmar

A Phlebotomidae sand fly, type species P. burmitis

Palaeomyia burmitis

Molluscs

Cephalopods

Name Novelty Status Authors Age Unit Location Notes Images
Conlinites[18]

Gen et sp nov

Valid

Kennedy

Early Cretaceous (Albian)

Paw Paw Formation

An ammonite, type species is C. wrighti.

Enigmaticeras[18]

Gen et sp nov

Valid

Kennedy

Early Cretaceous (Albian)

Paw Paw Formation

An ammonite, type species is E. riceae.

Ficheuria americana[18]

Sp nov

Valid

Kennedy

Early Cretaceous (Albian)

Paw Paw Formation

An ammonite.

Mariella (Mariella) asper[18]

Sp nov

Valid

Kennedy

Early Cretaceous (Albian)

Paw Paw Formation

An ammonite.

Neophlycticeras (Paradolphia) occidentalis[18]

Sp nov

Valid

Kennedy

Early Cretaceous (Albian)

Paw Paw Formation

An ammonite.

Newly named fishes

Name Novelty Status Authors Age Unit Location Notes Images

Gomphonchus? minutus[19]

Sp. nov

Valid

Valiukevičius

Late Silurian

Minija Formation

 Latvia

An acanthodian belonging to the group Ischnacanthiformes and the family Ischnacanthidae.

Lophosteus canadensis[20]

Sp. nov

Valid

Schultze & Märss

Late Silurian

Barlow Inlet Formation

 Canada

A bony fish belonging to the group Lophosteiformes.

Lophosteus ohesaarensis[20]

Sp. nov

Valid

Schultze & Märss

Late Silurian

 Estonia
 Latvia

A bony fish belonging to the group Lophosteiformes.

Lophosteus uralensis[20]

Sp. nov

Valid

Schultze & Märss

Early Devonian

 Russia

A bony fish belonging to the group Lophosteiformes.

Nostolepis alifera[19]

Sp. nov

Valid

Valiukevičius

Late Silurian

Targale Formation

 Latvia

An acanthodian belonging to the group Climatiiformes and the family Climatiidae.

Nostolepis latvica[19]

Sp. nov

Valid

Valiukevičius

Late Silurian

Minija Formation
Targale Formation

 Latvia

An acanthodian belonging to the group Climatiiformes and the family Climatiidae.

Perscheia[21]

Gen. et sp. nov

Valid

Elliott, Mark-Kurik & Daeschler

Devonian (Frasnian)

Nordstrand Point Formation

 Canada

A member of Pteraspidiformes belonging to the group Psammosteida and the family Obrucheviidae. The type species is P. pulla

Poracanthodes marssae[19]

Sp. nov

Valid

Valiukevičius

Late Silurian

Targale Formation

 Estonia
 Latvia

An acanthodian belonging to the group Ischnacanthiformes and the family Poracanthidae.

Poracanthodes sulcatus[19]

Sp. nov

Valid

Valiukevičius

Late Silurian

Targale Formation

 Estonia
 Latvia

An acanthodian belonging to the group Ischnacanthiformes and the family Poracanthidae.

Talimaalepis[22]

Gen. et sp. nov

Valid

Žigaitė

Silurian

Dashtygoi Formation
Kyzyl-Tchiraa Formation
Tchergak Formation

 Mongolia
 Russia

A thelodont, possibly a member of the group Phlebolepidiformes. The type species is T. rimae

Trimpleylepis[23]

Gen. et 2 sp. nov

Valid

Miller, Märss & Blom

Late Silurian

 Estonia
 United Kingdom

A member of Anaspida belonging to the group Birkeniida, possibly a member of the family Septentrioniidae. The type species is T. juncta; genus also includes T. concatenata.

Newly named amphibians

Name Novelty Status Authors Age Unit Location Notes Images

Jakubsonia[24]

Gen. et sp. nov

Valid

Lebedev

Devonian (Famennian)

 Russia

A basal member of Tetrapoda. The type species is J. livnensis

Reptile

Archosauromorphs

Newly named dinosaurs

Data courtesy of George Olshevsky's dinosaur genera list.[25]

Name Novelty Status Authors Age Unit Location Notes Images

Atrociraptor[26]

Valid

Currie & Varricchio

Edmontonian

Horseshoe Canyon Formation

 Canada

Atrociraptor

Bashunosaurus[27]

Gen et sp nov

Valid

Kuang

Middle Jurassic

Xiashaximiao Formation

 China

Bissektipelta[28]

Gen et sp nov

Valid

Parish & Barrett

Cenomanian-Turonian

Bissekty Formation

 Uzbekistan

Bonatitan[29]

Gen et sp nov

Valid

Martinelli & Forasiepi

Late Cretaceous

Bajo de Santa Rosa

 Argentina

Bonitasaura[30]

Gen et sp nov

Valid

Apesteguía

late Cretaceous

Bajo de la Carpa Formation

 Argentina

Bonitasaura

Borealosaurus[31]

Gen et sp nov

Valid

You, Ji, Lamanna,
Li, & Li

late Cretaceous

Sunjiawan Formation

 China

Dilong[32]

Gen et sp nov

Valid

Xu, Norell, Kuang,
Wang, Zhao, & Jia

Early Cretaceous

Yixian Formation

 China

Dilong

Ekrixinatosaurus[33]

Gen et sp nov

Valid

Calvo, Rubilar-Rogers, & Moreno

Late Cretaceous

Candeleros Formation

 Argentina

Ekrixinatosaurus

Graciliraptor[34]

Valid

Xing & Wang

Barremian

Yixian Formation

 China

Huaxiagnathus[35]

Valid

Hwang, Norell, Ji, & Gao

Barremian

Yixian Formation

 China

Huaxiagnathus

Kerberosaurus[36]

Valid

Bolotsky & Godefroit

Late Cretaceous

 Russia

Limaysaurus[37]

Valid

Salgado, Garrido, Cocca, & Cocca

 Argentina

Limaysaurus

Mei[38]

Valid

Xu & Norell

Barremian

Yixian Formation

 China

Mei

Mirischia[39]

Valid

Naish, Martill, & Frey

Albian

Santana Formation

 Brazil

Mirischia

Nemegtia[40]

Preoccupied name

Lü, Tomida, Azuma,
Dong, Lee, & Szczechura

Renamed Nemegtomaia

Otogosaurus

Nomen nudum

Zhao & Tan

Prenoceratops[41]

Valid

Chinnery

 USA
 Montana

Prenoceratops

Rugops[42]

Valid

Sereno, Wilson, & Conrad

 Niger

Rugops

Sinusonasus[43]

Valid

Xu, & Wang

Barremian

Yixian Formation

 China

Spinostropheus[42]

Valid

Sereno, Wilson & Conrad

Middle Jurassic

Irhazer Group

 Niger

A ceratosaur; new genus for
"Elaphrosaurus" gautieri Lapparent 1960.

Suuwassea[44]

Valid

Harris & Dodson

Kimmeridgian

Morrison Formation

 United States

Suuwassea

Talenkauen[45]

Valid

Novas, Cambiaso, & Ambrosio

Pari Aike Formation

 Argentina

Talenkauen

Tazoudasaurus[46]

Valid

Allain et al

 Morocco

Unaysaurus[47]

Valid

Leal, Azevedo,
Kellner, & Da Rosa

Caturrita Formation

 Brazil

Unaysaurus

Pterosaurs

New taxa
Name Status Authors Age Unit Location Notes

Avgodectes

Valid

Peters

Early Cretaceous

Yixian Formation

 China

A pterosaur hatchling.

Cacibupteryx

Valid

Gasparini Fernández de la Fuente

Late Jurassic (Oxfordian)

Jagua Formation

 Cuba

A rhamphorhynchid.

"Daitingopterus"

Nomen Nudum

Maisch Matzke Ge Sun

Late Jurassic (Tithonian)

Solnhofen Limestone

 Germany

A germanodactylid.

Lonchognathosaurus

Valid

Maisch, M.W., Matzke, A.T., and Ge Sun

Early Cretaceous (Aptian)

Lianmuqin Formation

 China

A dsungaripterid.

Other archosauromorphs

Name Status Authors Location Notes Images
Crosbysaurus[48] Valid non-dinosaurian taxon.  USA
Protecovasaurus[49] Valid non-dinosaurian taxon.
  • Heckert
 USA

Newly named birds

Name Status Novelty Authors Age Unit Location Notes Images

Aberratiodontus [50]

Valid

Gen et sp. nov.

Gong, Hou & Wang

Early Cretaceous

Jiufotang Formation

 China

An Enantiornithes,
Type species A. wui

Anserpica kiliani [51]

Valid

Gen et sp. nov.

Cécile Mourer-Chauviré

Didier Berthet

Marguerite Hugueney

Late Oligocene, Early Miocene

MP 29

 France

An Anseranatidae, the type species of Anserpica Mourer-Chauviré, Berthet & Hugueney, 2004.

Bavaripsitta ballmanni [52]

Valid

Gen et sp. nov.

Gerald Mayr

Ursula B. Göhlich

Middle Miocene

Mn 6

 Germany:

 Bavaria

A Psittacidae, the type species of Bavaripsitta Mayr & Göhlich, 2004.

Borvocarbo guilloti [51]

Valid

Gen et sp. nov.

Cécile Mourer-Chauviré

Didier Berthet

Marguerite Hugueney

Late Oligocene, Early Miocene

MP 30, MN 1

 France;

 Germany

A Phalacrocoracidae, the type species of Borvocarbo Mourer-Chauviré, Berthet & Hugueney, 2004.

?Colymboides metzleri [53]

Valid

Sp. nov.

Gerald Mayr

Early Oligocene

Rupelian

 Germany:

 Baden-Württemberg;

 Belgium:

 Antwerp

A Gaviidae, not certain a species of Colymboides Milne-Edwards, 1868.

Eurotrochilus inexpectatus [54]

Valid

Gen et sp. nov.

Gerald Mayr

Early Oligocene

Rupelian

 Germany:

 Baden-Württemberg

A Trochilidae, the type species of Eurotrochilus Mayr, 2004.

Iaceornis marshi [55]

Valid

Gen et sp. nov.

Julia A. Clarke

Late Cretaceous

Niobrara Formation

 USA:

 Kansas

An Ornithurae, the type species of Iaceornis Clarke, 2004.

Longirostravis hani [56]

Valid

Gen et sp. nov.

Hou Lianhai

Luis M. Chiappe

Zhang Fucheng

Chuong Cheng-Ming

Early Cretaceous

Yixian Formation

 China

An Enantiornithes, Euenantiornithes, the type species of Longirostravis Hou, Chiappe, Zhang et Chuong, 2004.

Meridiocichla salotti [57]

Valid

Gen et sp. nov.

Antoine Louchart

Late Quaternary

Corsica, Crete

 France:

 Corsica;

 Greece

A Turdidae, the type species of Meridiocichla Louchart, 2004.

Miopica paradoxa [58]

Valid

Gen et sp. nov.

Evgeny N. Kurochkin

Denis V. Sobolev

Miocene

Middle Meothis

MN 10

 Ukraine

A Corvidae, the type species of Miopica Kurochkin & Sobolev, 2004.

Mirolia brevirostrata [59]

Valid

Gen et sp. nov.

Peter Ballmann

Middle Miocene

MN 6

 Germany:

 Bavaria

A Scolopacidae, the type species of Mirolia Ballmann, 2004.

Mirolia dubia [59]

Valid

Sp. nov.

Peter Ballmann

Middle Miocene

MN 6

 Germany:

 Bavaria

A Scolopacidae.

?Mirolia mascalidris [59]

Valid

Sp. nov.

Peter Ballmann

Middle Miocene

MN 6

 Germany:

 Bavaria

A Scolopacidae, not certain to belong in Mirolia Ballmann, 2004.

Mirolia parvula [59]

Valid

Sp. nov.

Peter Ballmann

Middle Miocene

MN 6

 Germany:

 Bavaria

A Scolopacidae.

Primobucco frugilegus [60]

Valid

Sp. nov.

Gerald Mayr

Cécile Mourer-Chauviré

Ilka Weidig

Middle Eocene

Messel pit,

MP 11

 Germany:

 Hessen

A Primobucconidae Feduccia et Martin, 1976.

Primobucco perneri [60]

Valid

Sp. nov.

Gerald Mayr

Cécile Mourer-Chauviré

Ilka Weidig

Middle Eocene

Messel pit

MP 11

 Germany:

 Hessen

A Primobucconidae Feduccia et Martin, 1976.

Ramphastosula ramirezi [61]

Valid

Gen et sp. nov.

Marcelo Stucchi

Mario Urbina

Early Pliocene

Pisco Formation

 Peru

A Sulidae, the type species of Ramphastosula Stucchi & Urbina, 2004.

Tyto mourerchauvireae [62]

Valid

Sp. nov.

Marco Pavia

Early-Middle Pleistocene

Sicily

 Italy:

 Sicily

A Tytonidae.

Vescornis hebeiensis [63]

Valid

Gen et sp. nov.

Zhang Fucheng

Per G.P. Ericson

Zhou Zhonghe

Early Cretaceous

Yixian Formation

 China

An Enentiornithes Walker, 1981, this is the type species of the new genus, most probably a junior synonym of Hebeiornis fengningensis Yan, 1999.

Vitirallus watlingi [64]

Valid

Gen et sp. nov.

Trevor H. Worthy

Holocene

Viti Levu

 Fiji

A Rallidae, the type species of Vitirallus Worthy, 2004.

Wingegyps cartellei [65]

Valid

Gen et sp. nov.

Herculano M. F. de Alvarenga

Storrs L. Olson

Late Pleistocene or Early Holocene

Gruta dos Brejões

 Brazil

A Cathartidae, the type species of Wingegyps Alvarenga & Olson, 2004.

Plesiosaurs

New taxa

Name Status Authors Age Unit Location Notes
Plesiopterys Valid O'Keefe Toarcian Posidonienschiefer  Germany

Synapsids

Non-mammalian

Name Status Authors Age Unit Location Notes Images
Ianthodon[66] Valid Kissel & Reisz Kasimovian  USA
Lobalopex
Lobalopex[67] Valid Sidor, Hopson and Keyser Lopingian Teekloof Formation  South Africa
Progalesaurus[68] Valid Sidor & Smith Induan Balfour Formation  South Africa
Pyozia [69] Valid Anderson and Reisz Capitanian Krasnoschelsk Formation  Russia
Rewaconodon[70] Valid Datta, Das & Luo Late Triassic Tiki Formation  India

Other Animals

Name Status Authors Age Unit Location Notes Images
Charniodiscus procerus Valid LaFlamme et al., 2004 Ediacaran Mistaken Point Formation and Trepassey Formation  Canada

References

  1. ^ Gini-Newman, Garfield; Graham, Elizabeth (2001). Echoes from the past: world history to the 16th century. Toronto: McGraw-Hill Ryerson Ltd. ISBN 9780070887398. OCLC 46769716.
  2. ^ Poinar, G. (2008). "Lutzomyia adiketis sp. n. (Diptera: Phlebotomidae), a vector of Paleoleishmania neotropicum sp. n. (Kinetoplastida: Trypanosomatidae) in Dominican amber". Parasites & Vectors. 1 (1): 22. doi:10.1186/1756-3305-1-22. PMC 2491605. PMID 18627624.
  3. ^ a b Smith, S.Y.; Currah, R.S.; Stockey, R.A. (2004). "Cretaceous and Eocene poroid hymenophores from Vancouver Island, British Columbia". Mycologia. 96 (1): 180–186. doi:10.2307/3762001. JSTOR 3762001. PMID 21148842.
  4. ^ Manchester, S. R.; Dillhoff, R. M. (2004). "Fagus (Fagaceae) fruits, foliage, and pollen from the Middle Eocene of Pacific Northwestern North America". Canadian Journal of Botany. 82 (10): 1509–1517. doi:10.1139/b04-112.
  5. ^ Pigg, K. B.; Ickert-Bond, S. M.; Wen, J. (2004). "Anatomically preserved Liquidambar (Altingiaceae) from the middle Miocene of Yakima Canyon, Washington state, USA, and its biogeographic implications". American Journal of Botany. 91 (3): 499–509. doi:10.3732/ajb.91.3.499. PMID 21653405.
  6. ^ DeVore, M.L.; Moore, S.M.; Pigg, K.B.; Wehr, W.C. (2004). "Fossil Neviusia leaves (Rosaceae: Kerrieae) from the Lower Middle Eocene of Southern British Columbia". Rhodora. 12 (927): 197–209. JSTOR 23314752.
  7. ^ Manchester, S.R.; Pigg, K.B.; Crane, P.R. (2004). "Palaeocarpinus dakotensis sp. n. (Betulaceae: Coryloideae) and associated staminate catkins, pollen, and leaves from the Paleocene of North Dakota". International Journal of Plant Sciences. 165 (6): 1135–1148. doi:10.1086/423870.
  8. ^ Wunderlich, J. (2004). "Fossil spiders in amber and copal. Conclusions, revisions, new taxa and family diagnoses of fossil and extant taxa". Beiträge zur Araneologie. 3AB: 1–1908.
  9. ^ Penney, D.; Ono, H.; Selden, P.A. (2005). "A new synonymy for the Madagascan copal spider fauna (Araneae, Selenopidae)" (PDF). Journal of Afrotropical Zoology. 2: 41–44.[permanent dead link]
  10. ^ Bosselaers, J. (2004). "A new Garcorops species from Madagascar copal (Araneae: Selenopidae)" (PDF). Zootaxa. 445: 1–7. doi:10.11646/zootaxa.445.1.1. S2CID 47620450.
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  12. ^ a b c Koteja, J (2004). "Scale insects (Hemiptera: Coccinea) from cretaceous Myanmar (Burmese) amber". Journal of Systematic Palaeontology. 2 (2): 109–114. Bibcode:2004JSPal...2..109K. doi:10.1017/S1477201904001166. S2CID 86393735. – via Taylor & Francis (subscription required)
  13. ^ Fernández, F. (2004). "The American species of the myrmicine ant genus Carebara Westwood (Hymenoptera: Formicidae)". Caldasia. 26: 191–238.
  14. ^ Engel, M.S.; Ortega-Blanco, J. (2008). "The fossil crown wasp Electrostephanus petiolatus Brues in Baltic Amber (Hymenoptera, Stephanidae): designation of a neotype, revised classification, and a key to amber Stephanidae". ZooKeys (4): 55–64. Bibcode:2008ZooK....4...55E. doi:10.3897/zookeys.4.49. hdl:2445/36428.
  15. ^ Nel, A.; Perrault, G.; Perrichot, V.; Néradeau, D. (2004). "The oldest ant in the lower cretaceous amber of Charente-maritime (SW France) (Insecta: Hymenoptera: Formicidae)". Geologica Acta. 2 (1): 23–29.
  16. ^ Archibald, S.B.; Makarkin, V.N. (2004). "New genus of minute Berothidae (Neuroptera) from Early Eocene amber of British Columbia" (PDF). The Canadian Entomologist. 136: 61–76. CiteSeerX 10.1.1.552.2285. doi:10.4039/n03-043. S2CID 36459014. Archived from the original (PDF) on 2011-08-18. Retrieved 2010-04-09.
  17. ^ Poinar, G. O. Jr. (2004). "Palaeomyia burmitis (Diptera: Phlebotomidae), a new genus and species of Cretaceous sand flies with evidence of blood-sucking habits". Proceedings of the Entomological Society of Washington. 106: 598–605.
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  20. ^ a b c Schultze, Hans-Peter; Märss, Tiiu (2004). "Revisiting Lophosteus, a primitive osteichthyan" (PDF). Acta Universitatis Latviensis. Earth and Environment Sciences. 679: 57–78.
  21. ^ Elliott, David K.; Mark-Kurik, Elga; Daeschler, Edward B. (2004). "A revision of Obruchevia (Psammosteida: Heterostraci) and a description of a new obrucheviid from the Late Devonian of the Canadian Arctic" (PDF). Acta Universitatis Latviensis. Earth and Environment Sciences. 679: 22–45.
  22. ^ Žigaitė, Živilė (2004). "A new thelodont from Lower Silurian of Tuva and north-west Mongolia" (PDF). Acta Universitatis Latviensis. Earth and Environment Sciences. 679: 158–165.
  23. ^ Miller, C. Giles; Märss, Tiiu; Blom, Henning (2004). "New anaspid material from the Late Silurian of Britain and Estonia" (PDF). Acta Universitatis Latviensis. Earth and Environment Sciences. 679: 46–56.
  24. ^ Lebedev, Oleg A. (2004). "A new tetrapod Jakubsonia livnensis from the Early Famennian (Devonian) of Russia and palaeoecological remarks on the Late Devonian tetrapod habitats" (PDF). Acta Universitatis Latviensis. Earth and Environment Sciences. 679: 79–98.
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