^Dufresnes, C.; et al. Fifteen shades of green: The evolution of Bufotes toads revisited. Molecular Phylogenetics and Evolution. 2019, 141: 106615. PMID 31520778. doi:10.1016/j.ympev.2019.106615.
^Dufresnes, C. Amphibians of Europe, North Africa and the Middle East: A Photographic Guide. Bloomsbury Publishing. 2019: 71, 76–80. ISBN 978-1-4729-4137-4.
^Hopkins, G.R.; E.D. Brodie, Jr. Occurrence of Amphibians in Saline Habitats: A Review and Evolutionary Perspective. Herpetological Monographs. 2015, 29 (1): 1–27. doi:10.1655/HERPMONOGRAPHS-D-14-00006.