卷缘桩菇
皮埃尔·比利亚尔於1785年繪製的卷缘桩菇
卷缘桩菇 (學名 :Paxillus involutus )又稱卷边网褶菌 或卷边桩菇 ,最早於1785年由皮埃尔·比利亚尔 描述發表,當時被歸在傘菌屬 [ 3] ,但翌年由奥古斯特·巴奇 發表的學名Agaricus involutus 才被視作有效描述[ 1] [ 4] ;1838年埃利亚斯·芒努斯·弗里斯 定立新屬樁菇屬 ,並將本種作為模式種 [ 5] 。本種外型如棕色、木質的陀螺 ,子實體 可高約6公分[ 6] ,蕈傘 直徑為4—12公分,邊緣向下卷曲[ 7] ;菌髓 黃色且多汁,並具特殊氣味[ 7] [ 8] ;孢子印 為棕色,擔孢子 呈橢球形,長7.5—9微米,寬5—6微米[ 9] 。許多乳菇屬 物種、黑毛樁菇 與耳狀網褶菌 等蕈類的外形皆與本種相似[ 10] [ 11] ,另外同屬物種Paxillus obscurisporus 為自本種分出,其子實體較本種為大[ 12] 。
卷缘桩菇可與許多針葉樹 或落葉樹 形成外菌根 [ 13] ,常在外菌根相關研究中被用作模式生物 ,其基因組 已被完整定序 [ 14] 。有研究顯示本種與樹木共生時可增加樹木根部對土壤真菌鐮刀菌 感染的抗性[ 15] [ 16] ,並抑制植株吸收土壤中的有害重金屬 [ 17] [ 18] 。本種為北半球 溫帶地區常見的蕈類,分布範圍包括歐洲、亞洲與北美洲[ 10] [ 19] ,在印度 [ 20] 、伊朗 [ 21] 、中國 [ 22] 、日本 也有分布,最北可至阿拉斯加 與格陵蘭 [ 23] [ 24] ,並已作為外來種 被引入澳洲 [ 25] [ 26] 、紐西蘭 [ 27] 與南美洲 的智利 [ 28] 。二戰以前卷缘桩菇在中歐與東歐都被認為在烹煮過可食[ 7] [ 19] ,僅知生食後可能引起腸胃症狀[ 9] ,但1944年德國真菌學家尤利烏斯·舍費爾 食用此菌後中毒身亡[ 19] [ 29] ,後來研究發現卷缘桩菇的一個抗原 可激活自體免疫反應 ,使人體的免疫系統攻擊自身的紅血球 ,以致自體免疫性溶血 ,多發生在長期食用此菌者[ 9] [ 30] 。1990年代起本種逐漸被列為不可食用的蕈類[ 31] ,但東歐每年仍每年有人食用此菌後中毒而死[ 32] [ 33] [ 34] [ 35] 。
參考文獻
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Paxillus involutus Agaricus involutus