sister-group to all other ferns †Rhyniophyta—rhyniophytes †Zosterophyllophyta—zosterophylls †Trimerophytophyta—trimerophytes †Progymnospermophyta Seed plants (spermatophytes)...

†Cooksonia †Uskiella †Rhyniophytes Lycophytes Lycopsida-clubmosses †Zosterophylls Euphyllophytes †Eophyllophyton †Psilophyton – trimerophytes Moniliformopses...