National Academy of Sciences (
1980–) (Member of the National Academy of Sciences of the United States)
[6]American Academy of Arts and Sciences (
1984–)
[7]Academia Europaea (
2002–) (tilknytning: AE section Cell and developmental biology)
[8][9]American Society for Cell Biology (
2017–)
[10]American Society for Microbiology
American Society for Virology
Alpha Omega Alpha
Royal Society (
2008–) (utenlandsk medlem av Royal Society)
[11]