Deutsche Akademie der Naturforscher Leopoldina (
1991–)
Academia Europaea (
1989–) (tilknytning: AE section Cell and developmental biology)
[2]National Academy of Sciences (
1990–) (Foreign Associate of the National Academy of Sciences)
[3]Berlin-Brandenburgische Akademie der WissenschaftenNordrhein-Westfalens akademi for vitenskap og kunst
Heidelberger Akademie der Wissenschaften (
1989–) (korresponderende medlem)
[4]Det franske vitenskapsakademiet (
2009–) (utanlandsk medlem)
American Academy of Arts and Sciences
Göttingens vitenskapsakademi
European Molecular Biology Organization
Royal Society (
1990–) (utenlandsk medlem av Royal Society)
[5]