명백한 화석 기록은 트라이아스기까지 거슬러 올라가며, 이 딱정벌레의 기원은 중생대 초기에 시작된 것으로 추정하고 있다.[4]
각주
↑M.Hansen. Phylogeny and classification of the staphyliniform beetle families (Coleoptera). Biologiske Skrifter 48, Copenhagen, 1997
↑Beutel, R. G. and Leschen, R.A.B. 2005. Phylogenetic analysis of Staphyliniformia (Coleoptera) based on characters of larvae and adults. Systematic Entomology
↑J. F. Lawrence and A. F. Newton, Jr. 1995. Families and subfamilies of Coleoptera (with selected genera, notes, references and data on family-group names). pp. 779-1006 In: J. Pakaluk & S. A. Slipinski (Eds.): Biology, Phylogeny, and Classification of Coleoptera: Papers Celebrating the 80th Birthday of Roy A. Crowson. Museum i Instytut Zoologii PAN, Warszawa
↑Grimaldi, D. and Engel, M.S. (2005). Evolution of the Insects. Cambridge University Press