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Troctomorpha; Temporal range: Late Jurassic–Present PreꞒ Ꞓ O S D C P T J K Pg N
	Burmacompsocus perreaui (Compsocidae)
	Liposcelis (Liposcelididae)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Psocodea
Suborder:	Troctomorpha; Roesler, 1944
Infraorders
	Amphientometae Amphientomidae; Compsocidae; †Electrentomidae; Manicapsocidae; Musapsocidae; Protroctopsocidae; Troctopsocidae; ; Nanopsocetae Liposcelidae; Pachytroctidae; Sphaeropsocidae; Phthiraptera (lice); ;

Troctomorpha is one of the three major suborders of Psocodea (formerly Psocoptera)^[1]^[2](barklice, booklice, and parasitic lice), alongside Psocomorpha and Trogiomorpha. There are more than 30 families and 5,800 described species in Troctomorpha.^[3]^[4]^[1] The order includes parasitic lice, which are most closely related to the booklice family Liposcelididae.

Cladogram

Cladogram showing the position of Troctomorpha within Psocodea:^[5]

Fossil record

The oldest record of the suborder is suggested to be Paramesopsocus adibi, known from the Late Jurassic Karabastau Formation of Kazakhstan.^[6]^[7]

Classification

Troctomorpha contains the following subgroups:^[1]

Infraorder: Amphientometae
- Family: Amphientomidae Enderlein, 1903 (tropical barklice)
- Family: Compsocidae Mockford, 1967
- Family: Electrentomidae Enderlein, 1911
- Family: Manicapsocidae Mockford, 1967
- Family: Musapsocidae Mockford, 1967
- Family: Protroctopsocidae Smithers, 1972
- Family: Troctopsocidae Mockford, 1967
Infraorder: Nanopsocetae
- Family: Liposcelididae Broadhead, 1950 (booklice)
- Family: Pachytroctidae Enderlein, 1904 (thick barklice)
- Family: Sphaeropsocidae Menon, 1941
- Parvorder: Phthiraptera
  - Superfamily: Amblycera
    - Family: Ancistronidae
    - Family: Boopiidae Mjoberg, 1910
    - Family: Colpocephalidae Eichler, 1937
    - Family: Gliricolidae
    - Family: Gyropidae Kellogg, 1896
    - Family: Laemobothriidae Mjoberg, 1910
    - Family: Menoponidae Mjoberg, 1910 (chicken body lice)
    - Family: Pseudomenoponidae Mjoberg, 1910
    - Family: Ricinidae Neumann, 1890
    - Family: Somaphantidae Eichler, 1941
    - Family: Trimenoponidae
    - Family: Trinotonidae Eichler, 1941
  - Superfamily: Anoplura
    - Family: Echinophthiriidae Enderlein, 1904 (seal lice)
    - Family: Enderleinellidae Ewing, 1929
    - Family: Haematopinidae Enderlein, 1904 (ungulate lice)
    - Family: Hamophthiriidae Johnson, 1969
    - Family: Hoplopleuridae Ewing, 1929 (armoured lice)
    - Family: Hybothiridae Ewing, 1929
    - Family: Linognathidae Webb, 1946 (pale lice)
    - Family: Microthoraciidae Kim & Lugwig, 1978
    - Family: Neolinognathidae Fahrenholz, 1936
    - Family: Pecaroecidae Kéler, 1963
    - Family: Pedicinidae Enderlein, 1904
    - Family: Pediculidae Leach, 1817 (body lice, head lice)
    - Family: Polyplacidae Fahrenholz, 1912 (spiny rat lice)
    - Family: Pthiridae Ewing, 1929 (crab lice or pubic lice)
    - Family: Ratemiidae Kim & Lugwig, 1978
  - Superfamily: Ischnocera (paraphyletic)
    - Family: Bovicolidae
    - Family: Dasyonygidae
    - Family: Goniodidae
    - Family: Heptapsogasteridae
    - Family: Lipeuridae
    - Family: Philopteridae Nitzsch, 1818 (paraphyletic)
    - Family: Trichodectidae
    - Family: Trichophilopteridae
  - Superfamily: Rhyncophthirina
    - Family: Haematomyzidae Enderlein, 1904

References

^ ^a ^b ^c ^d Johnson, Kevin P.; Smith, Vincent S. (2021). "Psocodea species file online, Version 5.0". Retrieved 2021-11-05.
^ Bess, Emilie; Smith, Vince; Lienhard, Charles; Johnson, Kevin P. (2006). "Psocodea". Tree of Life. Retrieved 2019-06-14.
^ "Troctomorpha Report". Integrated Taxonomic Information System. Retrieved 2019-05-04.
^ "Troctomorpha suborder Information". BugGuide.net. Retrieved 2019-05-04.
^ de Moya, Robert S; Yoshizawa, Kazunori; Walden, Kimberly K O; Sweet, Andrew D; Dietrich, Christopher H; Kevin P, Johnson (2021-06-16). Buckley, Thomas (ed.). "Phylogenomics of Parasitic and Nonparasitic Lice (Insecta: Psocodea): Combining Sequence Data and Exploring Compositional Bias Solutions in Next Generation Data Sets". Systematic Biology. 70 (4): 719–738. doi:10.1093/sysbio/syaa075. ISSN 1063-5157. PMID 32979270.
^ Azar, Dany; Hajar, Lara; Indary, Chadi; Nel, André (January 2008). "Paramesopsocidae, a new Mesozoic psocid family (Insecta: Psocodea "Psocoptera": Psocomorpha)". Annales de la Société entomologique de France. New Series. 44 (4): 459–470. doi:10.1080/00379271.2008.10697581. ISSN 0037-9271. S2CID 85044414.
^ Mockford, Edward L.; Lienhard, Charles; Yoshizawa, Kazunori (October 2013). "Revised classification of 'Psocoptera' from Cretaceous amber, a reassessment of published information". Insecta Matsumurana. New Series. 69: 1–26. ISSN 0020-1804.

Troctomorpha

Cladogram

Fossil record

Classification

References

Further reading