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Thoracotremata is a clade of crabs, comprising those crabs in which the genital openings in both sexes are on the sternum, rather than on the legs. It comprises 17 families in four superfamilies .[1]
Thoracotremata is the sister group to Heterotremata within the clade Eubrachyura, having diverged during the Cretaceous period. Eubrachyura itself is a subset of the larger clade Brachyura, which consists of all "true crabs". A summary of the high-level internal relationships within Brachyura can be shown in the cladogram below: [2][3]
Dromiacea
Raninoida
Cyclodorippoida
Heterotremata
Thoracotremata
The internal relationships within Thoracotremata are less certain, with many of the superfamilies found to be invalid. The proposed cladogram below is from analysis by Tsang et al, 2014:[2]
Pinnotheridae
Dotillidae
Percnidae
Xenograpsidae
Cryptochiridae
Ocypodidae
Glyptograpsidae
Grapsidae
Plagusiidae
Gecarcinidae
Sesarmidae
Mictyridae
Varunidae
Macrophthalmidae
However, recent studies have found the superfamilies Grapsoidea and Ocypodoidea to be polyphyletic and invalid.[2][3]
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