During World War II, Tarzan reverts to his identity of John Clayton, Lord Greystoke, and resumes his military career as a pilot for the RAF. Flying a P-40 Tomahawk, Clayton is sent to rescue a missing British Military Intelligence officer, code-named Ilex. When his plane is brought down by a pteranodon over Pal-ul-don, his adventure returns him to the lost land, where he befriends the elephant Torn Ear and confronts the mysterious Turtle People. Eventually he even finds and recovers Ilex.