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Strombus gracilior; Temporal range: Pleistocene–Recent PreꞒ Ꞓ O S D C P T J K Pg N ↓
	Apertural view of a shell of the Eastern Pacific fighting conch, Strombus gracilior
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Subclass:	Caenogastropoda
Order:	Littorinimorpha
Family:	Strombidae
Genus:	Strombus
Species:	S. gracilior
Binomial name
	Strombus gracilior; Sowerby I, 1925

Strombus gracilior, common names the Eastern Pacific fighting conch, or the Panama fighting conch, is a species of medium to large sea snail, a marine gastropod mollusk in the family Strombidae, the true conchs.^[1]

Description

The length of the shell varies between 40 mm and 95 mm, the width may attain 50 mm. The high spire is covered with subsutural spines or pointed nodes on the shoulder of the whorls. The color of the shell is yellowish to yellowish-brown, interrupted in the middle with a lighter band. The aperture and the large outer lip is white bordered with orange-brown. The shell is covered with a thin, horn-covered periostracum.

The fighting conch (Strombus pugilis) from the Caribbean Sea and the Gulf of Mexico is a similar species in the sculpture of the shell and in inner morphological characters.^[2]

Phylogeny

Strombidae

Terebellum terebellum

	Eustrombus

	Aliger

A simplified version of the phylogeny and relationships of Strombidae according to Simone (2005)^[2]

	Strombus gallus

	Strombus gigas

Strombus costatus

	Strombus raninus

	Strombus peruvianus

Strombus galeus

Strombus latus

	Strombus pugilis

	Strombus alatus

Strombus gracilior

Strombus granulatus

Phylogeny and relationships of Eastern Pacific and Atlantic Strombus species, according to Latiolais et al. (2006)^[3]

The phylogenetic relationships among the Strombidae have been mainly accessed on two occasions, using two distinct methods. In a 2005 monography, Simone proposed a cladogram (a tree of descent) based on an extensive morpho-anatomical analysis of representatives of Aporrhaidae, Strombidae, Xenophoridae and Struthiolariidae.^[2] However, according to Simone, only Strombus gracilior, Strombus alatus and Strombus pugilis, the type species, remained within Strombus. In Simone's cladogram, these three species constituted a distinct group based on at least five synapomorphies (traits that are shared by two or more taxa and their most recent common ancestor). The remaining taxa were previously considered as subgenera, and were elevated to genus level by Simone in the end of his analysis.^[2]

In a different approach, Latiolais and colleagues (2006) proposed another cladogram that attempts to show the phylogenetic relationships of 34 species within the family Strombidae. The authors analysed 31 species in the genus Strombus and three species in the allied genus Lambis. The cladogram was based on DNA sequences of both nuclear histone H3 and mitochondrial cytochrome-c oxidase I protein-coding gene regions. In this proposed phylogeny, Strombus pugilis, Strombus alatus, Strombus granulatus and Strombus gracilior are closely related and appear to share a common ancestor.^[3]

Distribution

This species is can be found on sandflats and lagoons and offshore to 45 m in the Gulf of California along West Mexico and in the Pacific Ocean along Northern Peru.