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Spialia
	Spialia mafa in the Ngorongoro Crater, Tanzania
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Lepidoptera
Family:	Hesperiidae
Tribe:	Carcharodini
Genus:	Spialia; Swinhoe, 1912

Spialia is a genus of skippers in the family Hesperiidae, which are mainly found in Africa and Asia. They are commonly called grizzled skippers or sandmen.

As a result of genomic research published in 2020, 3 species of Spialia were moved to the new genus Agyllia and 14 species were moved to the new genus Ernsta.^[1]

Species

These species belong to the genus Spialia:^[1]^[2]^[3]

Spialia ali Oberthür, 1881
Spialia carnea (Reverdin, 1927)
Spialia diomus (Hopffer, 1855) (common sandman)
Spialia doris (Walker, 1870) (desert grizzled skipper)
Spialia ferax (Wallengren, 1863) (ferax grizzled skipper)
Spialia fetida Zhdanko, 1992
Spialia galba (Fabricius, 1793) (indian skipper)
Spialia geron (Watson, 1893)
Spialia irida Zhdanko, 1993
Spialia lugens (Staudinger, 1886)
Spialia mafa (Trimen, 1870) (mafa sandman)
Spialia orbifer (Hübner, 1823) (orbed red-underwing skipper)
Spialia osthelderi (Pfeiffer, 1932)
Spialia phlomidis (Herrich-Schäffer, 1845) (persian skipper)
Spialia rosae Hernández-Roldán, Dapporto, Dinca, Vicente & Vila, 2016
Spialia sertorius (Hoffmannsegg, 1804) (red-underwing skipper)
Spialia spio (Linnaeus, 1764) (mountain sandman)
Spialia struvei (Püngeler, 1914)
Spialia therapne (Rambur, 1832) (corsican red-underwing skipper)

References

^ ^a ^b Zhang, Jing; Brockmann, Ernst; Cong, Qian; Shen, Jinhui; et al. (2020). "A genomic perspective on the taxonomy of the subtribe Carcharodina (Lepidoptera: Hesperiidae: Carcharodini)". Zootaxa. 4748 (1): zootaxa.4748.1.10. doi:10.11646/zootaxa.4748.1.10. PMC 8018707. PMID 32230093.
^ Spialia at Markku Savela's Lepidoptera and Some Other Life Forms
^ Hernández-Roldán, Juan L.; Dapporto, Leonardo; Dincă, Vlad; Vicente, Juan C.; Hornett, Emily A.; Šíchová, Jindra; Lukhtanov, Vladimir; Talavera, Gerard; Vila, Roger (2016). "Integrative analyses unveil speciation linked to host plant shift in Spialia butterflies". Molecular Ecology. 25 (17): 4267–84. Bibcode:2016MolEc..25.4267H. doi:10.1111/mec.13756. hdl:10230/32118. PMID 27393640.

Natural History Museum Lepidoptera genus database
Woodhall, Steve (2005). Field Guide to Butterflies of South Africa. Cape Town, South Africa: Struik. ISBN 978-1-86872-724-7.
Seitz, A. Die Gross-Schmetterlinge der Erde 13: Die Afrikanischen Tagfalter. Plate XIII 75

External links

Images representing Spialia at Consortium for the Barcode of Life

This Pyrginae-related article is a stub. You can help Wikipedia by expanding it.

[Zhang2020-1] Zhang, Jing; Brockmann, Ernst; Cong, Qian; Shen, Jinhui; et al. (2020). "A genomic perspective on the taxonomy of the subtribe Carcharodina (Lepidoptera: Hesperiidae: Carcharodini)". Zootaxa. 4748 (1): zootaxa.4748.1.10. doi:10.11646/zootaxa.4748.1.10. PMC 8018707. PMID 32230093.

[funet-2] Spialia at Markku Savela's Lepidoptera and Some Other Life Forms

[Roldán2016-3] Hernández-Roldán, Juan L.; Dapporto, Leonardo; Dincă, Vlad; Vicente, Juan C.; Hornett, Emily A.; Šíchová, Jindra; Lukhtanov, Vladimir; Talavera, Gerard; Vila, Roger (2016). "Integrative analyses unveil speciation linked to host plant shift in Spialia butterflies". Molecular Ecology. 25 (17): 4267–84. Bibcode:2016MolEc..25.4267H. doi:10.1111/mec.13756. hdl:10230/32118. PMID 27393640.

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