Sanjuansaurus was named and described in 2010 by Oscar Alcober and Ricardo Martinez. The type species was named S. gordilloi after Raul Gordillo, the head fossil preparator and artist of the San Juan Museum. It is known from and based on an associated and partially articulated partial skeleton (PVSJ 605) consisting of a jaw fragment, most of the vertebral column from the axis to the anterior tail, the shoulder blades, an ulna, part of the pelvis, most of the long bones of the legs, and a few other bones.[1]
PVSJ 605 was discovered in 1994, in gray-green sandstone 40 meters above the base of the Ischigualasto Formation, in Ischigualasto Provincial Park in San Juan, Argentina. An ash bed from the early part of the formation dates to approximately 231.4 Ma, during the late Carnian Stage of the Late Triassic.[1] The original description indicated that the skeleton was from the earliest part of the Cancha de Bochas Member,[1] though later sources suggested that it was from the upper La Peña member.[2][3]
Description and classification
Sanjuansaurus was comparable in size to a medium-sized Herrerasaurus, with a thigh bone that was 395 millimetres (15.6 in) long and a tibia that is 360 millimetres (14 in) in length (in the description paper the tibia is listed with 260 mm in length due to a typo).[1] Alcober and Martinez performed a phylogenetic analysis and found Sanjuansaurus to be a herrerasaurid.[1] It was determined that Sanjuansaurus andHerrerasaurus share many similarities in the morphology of the skull, neck vertebrae, back vertebrae, hip vertebrae, scapula, and the hip bones. Alcober and Martinez observed that Sanjuansaurus andStaurikosaurusshare many similarities in the morphology of the hip bones, and the tibia.[1] The pubis of Sanjuansaurus, unlike in other herrerasaurids, points toward the cranium.
Distinguishing anatomical features
A diagnosis is a statement of the anatomical features of an organism (or group) that collectively distinguish it from all other organisms. Some, but not all, of the features in a diagnosis are also autapomorphies. An autapomorphy is a distinctive anatomical feature that is unique to a given organism or group.
According to Alcober and Martinez (2010), Sanjuansaurus can be distinguished based on the following characteristics:[1]
the cervical vertebrae have shelf-like, posterolaterally directed transverse processes
the neural spines of the sixth to eighth dorsal vertebrae, at least, bear acute anterior and posterior processes
the pubis is relatively short, measuring 63% of the length of the femur
a pronounced, rugose scar is present on the medial surface of the femur at the level of the fourth trochanter
Paleoecology
Fauna and habitat
In the Ischigualasto Formation, dinosaurs constituted only about 6% of the total number of fossils,[4] but by the end of the Triassic Period, dinosaurs were becoming the dominant large land animals, and the other archosaurs and synapsids declined in variety and number.[5]
^Martínez, Ricardo N.; Apaldetti, Cecilia; Alcober, Oscar A.; Colombi, Carina E.; Sereno, Paul C.; Fernandez, Eliana; Malnis, Paula Santi; Correa, Gustavo A.; Abelin, Diego (2013). "Vertebrate succession in the Ischigualasto Formation". Journal of Vertebrate Paleontology. 32 (Memoir 12: Basal sauropodomorphs and the vertebrate fossil record of the Ischigualasto Formation (Late Triassic: Carnian-Norian) of Argentina, sup1): 10–30. doi:10.1080/02724634.2013.818546. hdl:11336/7771. ISSN0272-4634. S2CID37918101.
^Benton, Michael J. (1999). "Origin and early evolution of dinosaurs". In Farlow, James O.; Brett-Surman, M.K. (eds.). The Complete Dinosaur. Indiana University Press. pp. 204–215. ISBN0-253-21313-4.
^Parrish, J. Michael (1999). "Evolution of the archosaurs". In Farlow, James O.; Brett-Surman, M.K. (eds.). The Complete Dinosaur. Indiana University Press. pp. 191–203. ISBN0-253-21313-4.
^Sereno, P.C. & Novas, F.E. (1992). The complete Skull and Skeleton of an Early dinosaur. Science. 258: 1137-1140. [Herrerasaurus ischigualastensis]
^Sill, W.D. (1974). "The anatomy of Saurosuchus galilei and the relationships of the rauisuchid thecodonts". Bulletin of the Museum of Comparative Zoology. 146: 317–362.
^Bonaparte, J.F. (1970). "Annotated list of the South American Triassic tetrapods". Gondwana Symposium Proceedings and Papers. 2: 665–682.