The ST staple is a common four- or five-amino acid residue motif in proteins and polypeptides with serine or threonine as the C-terminal residue.[1][2] It is characterized by a single hydrogen bond between the hydroxyl group of the serine or threonine (at residue i + 3 or i + 4) and the main chain carbonyl group of residue i. Motifs are of two types, depending whether the motif has 4 or 5 residues. Most ST staples occur in conjunction with an alpha helix, and are usually associated with a slight bend in the helix. Two websites are available for finding and examining ST staples in proteins: Motivated Proteins[3] and PDBeMotif.[4]
References
^Gray, TM; Matthews BW (1984). "Intrahelical hydrogen bonding of serine, threonine and cysteine residues within α-helices. Relevance to membrane-bound proteins". Journal of Molecular Biology. 175 (1): 75–82. doi:10.1016/0022-2836(84)90446-7. PMID6427470.