Though a majority of the taxa listed below lived during the latest early Campanian (c. 80.5 million years ago; fossils from the site Ivö Klack alone from this time compromise about 40 vertebrate species and more than 200 invertebrate species),[1] the Kristianstad Basin preserves fossils ranging in age from the early Santonian (c. 86.3 million years ago) to the early Maastrichtian (c. 72.1 million years ago); some of the animals in the list were not contemporaries, but separated from each other in time by several million years. The time spans from which fossils have been recovered is included for each species in the list.
A pachycormid fish. The teeth of Protosphyraena are much larger than those of the other fish in the basin and were once mistakenly referred to theropod dinosaurs.
Squalus in the Kristianstad Basin are from a younger (Maastrichtian) layer than the other sharks. Same genus as modern spurdogs. The Ballingslöv site is not part of the Kristianstad Basin proper, representing a northwestern isolated outcrop.
In addition to the remains referred to Aigialosuchus, detached and unidentified crocodylomorph scutes have also been discovered in Campanian-age deposits at Ivö Klack.[29]
A marine dyrosaurid crocodylomorph and the only crocodylomorph known from the Kristianstad Basin. Likely durophagous (eating hard-shelled organisms). Its skull was similar to that of modern gharials.
Four maxillary teeth, a dentary tooth, two caudal vertebrae, one right manual phalanx
Morphology of the teeth recovered corresponds roughly to Leptoceratops. Given that the teeth lack some of the ridges present in the teeth of Leptoceratops, and because of geographical distance, the Åsen leptoceratopsid is likely a new taxon.
Identified as the remains of a small theropod dinosaur after comparisons with the same bone in the Australian genus Australovenator. The bone structure confidently identifies the fossil as that of a non-avian theropod dinosaur.
Most of the Clidastes fossils in the Kristianstad Basin are probably from juveniles measuring about 2–3 meters in length, with a few adult individuals at probably around 6 meters.
Known from a single tooth, Mosasaurus in the Kristianstad Basin is from a younger (Late Campanian/Maastrichtian) layer than the other mosasaurs. Similar in body size (and likely ecological niche) to the earlier Tylosaurus ivoensis.
The overall morphology of the quadrate bone is consistent with Plioplatecarpus and in particular with P. primaevus, a species otherwise only found in North America. The quadrate is from a younger (Maastrichtian) layer than the other mosasaurs.
A mosasaur at about 8 meters in length with relatively wide and deep jaws. Bite marks matching Prognathodon teeth have been found on plesiosaur bones in the Kristianstad Basin. Frequently treated and listed under the name "Dollosaurus" or "Prognathodon (Dollosaurus)"; Dollosaurus being a junior synonym of Prognathodon.
Several marginal teeth, pterygoid teeth, partial jawbones and several vertebrae
Previously referred to Mosasaurus and Hainosaurus. The largest mosasaur in the basin and "undoubtedly" its top predator, the lower jaw of Tylosaurus ivoensis is estimated to have been over 1.5 meters in length.
The last comprehensive review of the plesiosaur fauna in the Kristianstad Basin was done by paleontologist Per-Ove Persson in the 1960s and his taxonomy is still used with caution, pending a much-needed new review.[38]
Vertebral centra, detached teeth, diaphysical ossifications of humeri and femora
The most common plesiosaur in the Kristianstad Basin. Scanisaurus is no longer considered a valid taxon, but the name continues to be used for practical purposes. S. nazarowi is the type species of Scanisaurus and was previously referred to Cimoliasaurus.
Fossils of this species include the only plesiosaurian skull (albeit crushed) known from Sweden. The preserved teeth are similar to those referred to Scanisaurus cf. nazarowi.
Possible pterosaur bone fragments have been recovered from earliest Campanian-age deposits at Ullstorp, though they remain unpublished.[8]
Scincomorpha
Fossils of terrestrial scincomorph lizards have been recovered in the Kristianstad Basin, but are as of yet unpublished.[59]
Turtles
In addition to the fossils described below, indeterminate turtle remains, including limb bones and carapace fragments, have also been recovered from Ivö Klack, Ugnsmunnarna, Ignaberga and Åsen.[10][60]
A macrobaenid turtle; Einarsson (2018) notes that the identification as Osteopygis is likely incorrect and that the material is more likely to belong to the genus Euclastes, though it has not formally been reclassified.
Note: Although the Kristianstad Basin is incredibly rich in invertebrate fossils and diversity, with hundreds of species, the creation of a full list is impossible due to a lack of published overviews and recent examinations.[63] Which lists are incomplete is specified below and estimates in regards to how many species are actually present are included if possible.
Bivalves
The bivalves are the most species-rich group present in the Kristianstad Basin, with Surlyk & Sørenson (2010) stating that close to 70 distinct species were present at Ivö Klack alone.[64]
Fossils of corals are common in several sites throughout the Kristianstad Basin.[76] The list below only accounts for the recently revised diversity of corals found at Ivö Klack.[86]
According to Surlyk & Sørenson (2010), 19 species of gastropods could be identified from fossils just from the latest Early Campanian of Ivö Klack.[64] Their subsequent 2011 study on the gastropods of the site only listed the 15 species accounted for below.[101]
In addition to the diverse polychaete worm fauna of Ivö Klack listed below, encrusters of serpulid polychaetes have also been discovered at Åsen,[11] though they are considerably fewer in number there and as of yet unpublished.[102]
Fossil sponges have been recovered at fossil sites in the Kristianstad Basin.[75]
Starfish
Fossils of starfish have been found at several sites in the Kristianstad Basin.[67] According to Surlyk & Sørenson (2010) there were 16 distinct species of starfish present at Ivö Klack alone.[64]
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