Hulley completed his MSc in 1967 from the University of Cape Town with a thesis entitled: Studies on the anatomy of some South African Mytilidae (Bivalvia) with notes on their ecology and distribution. He submitted his PhD thesis in 1971 entitled: The origin, interrelationship and distribution of Southern African Rajidae (Chondrichthyes, Batoidei)
He has worked at the South African Museum since 1965, where he was deputy director (research) and curator of fishes at the South African Museum until 2005. From 2006, Hulley has worked at the South African Museum in an honorary capacity as a research associate.[4]
The comprehensive Mesopelagic Fish Collection of the South African museum was assembled primarily by Hulley. The collection contains fish that live between 200m and 1 000m below the surface. Mesopelagic fish constitutes 95% of the world’s fish biomass.[5]
Selected publications
Hulley has authored or co-authored more than 100 articles, reports and chapters for academic works, including:
Polidoro, Beth A.; Ralph, Gina M.; Strongin, Kyle; Harvey, Michael; Hulley, Percival A.; et al. (15 February 2017). "The status of marine biodiversity in the Eastern Central Atlantic (West and Central Africa)". Aquatic Conservation: Marine and Freshwater Ecosystems. 27 (5). Wiley: 1021–1034. Bibcode:2017ACMFE..27.1021P. doi:10.1002/aqc.2744. ISSN1052-7613.
Duhamel, Guy; Belchier, Mark; Hulley, P. Alexander; et al. (January 2014). "BIOGEOGRAPHIC PATTERNS OF FISH. THE BIOGEOGRAPHIC ATLAS OF THE SOUTHERN OCEAN". doi:10.13140/2.1.1828.3203. {{cite journal}}: Cite journal requires |journal= (help)
^Pietsch, Theodore W.; Anderson, William Dewey (1997). Collection building in ichthyology and herpetology. American Society of Ichthyologists and Herpetologists. p. 129. ISBN9780935868913. Retrieved 9 July 2011. Accompanied by Stehmann and P. Alexander Hulley (Table 1), Krefft used the opportunity to complement his former Atlantic midwater transects by conducting two more transects along 50° W and 40° W from the latitude of Montevideo to the ...