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Nathrenus
	Anthrenus verbasci, common species from the subgenus (cosmopolitan)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Family:	Dermestidae
Genus:	Anthrenus
Subgenus:	Nathrenus; Casey, 1900
Species
	See text.

Nathrenus is a subgenus of the genus Anthrenus of the subfamily Megatominae within the family of skin beetles.^[1] The most well-known species from the subgenus, varied carpet beetle (Anthrenus verbasci), is distributed in most parts of the world, whereas the vast majority of other species are only present in regions of Africa and Asia.

Key characters

Subgenus is distinguished by antennae with 11 segments. Occasionally, male specimen appear with 10 antennae segments, having one less "filler" segment in the middle. Inner edge of eye doesn't have indentation, as opposed to subgenus Anthrenus.^[2]

Species

According to World Dermestidae catalogue, these species currently belong to subgenus Nathrenus:^[3]^[4]

Species group "albomaculatus"

Anthrenus albomaculatus (Pic, 1927) – Indonesia (Borneo, Kalimantan); Malaysia (Pahang, Perak)
Anthrenus cardamom (Háva, 2001) – South India
Anthrenus constantini (Háva & Herrmann, 2006) – South Africa
Anthrenus herrmanni (Kadej & Háva, 2016) – Thailand
Anthrenus hrdlickai (Háva, 2016) – Philippines (Luzon)
Anthrenus kalimantanus (Háva, 2004) – Indonesia (Kalimantan)
Anthrenus mindanaoensis (Háva, 2004) – Philippines
Anthrenus perak (Háva, 2016) – Indonesia (Kalimantan); Malaysia
Anthrenus preissi (Háva & Herrmann, 2003) – Indonesia (Kalimantan, Sumatra)
Anthrenus sabahense (Háva, 2021) – Malaysia (Sabah)
Anthrenus viktorai (Háva, 2016) – Malaysia

Species group "verbasci"

Anthrenus afer (Péringuey, 1886) – South Africa
Anthrenus albostictus (Reitter, 1881) – South Africa
Anthrenus arndti (Háva, 2005) – Namibia
Anthrenus barclayi (Háva, 2019) – Angola
Anthrenus basilewskyi (Kalík, 1965) – Kenya, Malawi, Tanzania, Zambia
Anthrenus bezdeki (Háva, 2017) – Yemen (Socotra)
Anthrenus biskrensis (Reitter, 1887) – Italy (Lampedusa, Lipari, Sicily); Malta; Spain; Algeria; Libya; Morocco; Tunisia
Anthrenus bomiensis (Háva, 2004) – China (Tibet, Yunnan)
Anthrenus boyesi (Háva, 2004) – Congo; Malawi; Swaziland; Zimbabwe; South Africa (Cape, Namaqualand, Natal, Transvaal, Zululand);
Anthrenus bulirschi (Háva, 2000) – Greece; Turkey; Israel; Jordan; Lebanon; Syria
Anthrenus capensis (Guérin–Méneville, 1835) – South Africa
Anthrenus cordis (Háva & Kadej, 2006) – South Africa
Anthrenus cylindricornis (Herrmann & Háva, 2014) – South Africa
Anthrenus edopetri (Háva, 2004) – Laos; Thailand
Anthrenus exilis (Mulsant & Rey, 1868) – Algeria; Egypt; Libya; Morocco; Tchad; Tunisia
Anthrenus funebris (Reitter, 1889) – Greece (Karpathos Islands)
Anthrenus gorki (Háva, 2008) – Azerbaijan; Greece; Turkey
Anthrenus havai (Kadej & Jakubska, 2007) – Namibia
Anthrenus jakli (Háva, 2001) – Oman; Yemen
Anthrenus kabakovi (Kadej & Háva, 2016) – Vietnam
Anthrenus knizeki (Háva, 2004) – China (Gansu, Hebei, Shaanxi, Sichuan)
Anthrenus kopeckyi (Háva, 2017) – Morocco
Anthrenus kucerai (Háva, 2005) – China (Sichuan)
Anthrenus liliputianus (Mulsant & Rey, 1868) – Egypt
Anthrenus longisetosus (Kadej & Háva, 2015) – China (Jiangxi, Yunnan, Zhejiang)
Anthrenus malawicus (Háva, 2004) – Malawi
Anthrenus maltzi (Kadej, 2010) – Angola
Anthrenus margarethae (Háva & Kadej, 2006) – South Africa
Anthrenus molitor (Aubé, 1850) – Albania; Bulgaria; Croatia; France; Greece; Italy; Macedonia; Spain; Turkey; Algeria; Egypt; Morocco; Tunisia; Iraq; Israel; Lebanon
Anthrenus nadeini (Kadej & Háva, 2008) – Ethiopia
Anthrenus narani (Háva & Ahmed, 2014) – Pakistan
Anthrenus natalensis (Háva, 2004) – South Africa (Cape, Natal)
Anthrenus nideki (Háva, 2005) – South Africa
Anthrenus nocivus (Mulsant & Godart, 1870) – Algeria
Anthrenus noctua (Háva, 2005) – South Africa
Anthrenus obenbergeri (Háva & Herrmann, 2021) – Crete
Anthrenus obscurus (Thunberg, 1815) – South Africa
Anthrenus paramolitor (Herrmann & Háva, 2021) – Crete; Turkey
Anthrenus pilosus (Pic, 1923) – China (Tibet)
Anthrenus propinquus (Háva, 2005) – China (Sichuan)
Anthrenus pubifer (Reitter, 1899) – Turkey; Iran; Iraq; Syria
Anthrenus purcharti (Háva, 2014) – Yemen (Socotra)
Anthrenus schawalleri (Háva & Kadej, 2006) – China (Sichuan)
Anthrenus sichuanicus (Háva, 2004) – China (Sichuan)
Anthrenus signatus (Erichson, 1846) – Austria; Bosnia & Herzegovina; Bulgaria; Croatia; Czech Republic; Greece; Hungary; Italy; Montenegro; Poland; Romania; Serbia; Slovakia; Slovenia; Spain; Turkey
Anthrenus snizeki (Háva, 2004) – South Africa
Anthrenus stelma (Kadej & Háva, 2006) – Namibia
Anthrenus strakai (Herrmann & Háva, 2012) – Cape Verde
Anthrenus subsetosus (Arrow, 1915) – Myanmar; Vietnam
Anthrenus transcaspicus (Mroczkowski, 1960) – Iran; Turkmenistan
Anthrenus tryznai (Háva, 2001) – China (Tibet)
Anthrenus turnai (Háva, 2004) – China (Sichuan)
Anthrenus undatus (Reitter, 1881) – South Africa
Anthrenus verbasci (Linnaeus, 1767) – Cosmopolitan
Anthrenus versicolor (Reitter, 1887) – Greece; Italy (Sardinia, Sicily)
Anthrenus zahradniki (Háva, 2003) – Greece (Chios, Rhodes); Lebanon; Turkey; Cyprus

Species group "ambericus" (extinct)

Some species of subgenus Nathrenus were preserved in amber and consist of:

†Anthrenus ambericus (Háva, Prokop & Herrmann, 2006)
†Anthrenus electron (Háva, Prokop & Kadej, 2006)
†Anthrenus groehni (Háva, Prokop & Herrmann, 2006)
†Anthrenus kerneggeri (Háva, Prokop & Herrmann, 2008)

Synonymization of Neoanthrenus

In 2013, researchers reclassified the species of the endemic Australian genus Neoanthrenus, placing them under the subgenus Nathrenus (as a group of species "ocellifer").^[5] However, recent research removes the species from genus Anthrenus due to significant genetic and morphological differences.^[6]

Species considered to be part of species group "ocellifer":^[3]

Anthrenus armstrongielus (Kadej & Háva, 2013) – Australia: Sydney
Anthrenus consobrinus (Háva, 2005) – Australia: Northern Territory, Western Australia
Anthrenus frater (Arrow, 1915) – Tasmania
Anthrenus king (Háva, 2002) – Australia (King Island)
Anthrenus macqueeni (Armstrong, 1949) – Australia (Queensland)
Anthrenus niveosparsus (Armstrong, 1941) – Australia (New South Wales)
Anthrenus ocellifer (Blackburn, 1891) – Australia (New South Wales, South Australia, Western Australia)
Anthrenus parallelus (Armstrong, 1941) – Australia (New South Wales, Queensland, Victoria)
Anthrenus svatopluki (Kadej & Háva, 2013) – Australia (Western Australia)