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Icla Formation

Icla Formation
Stratigraphic range: Pragian - Emsian
TypeGeological formation
UnderliesHuamampampa Formation
OverliesSanta Rosa Formation
Location
RegionSub-alpine region [es]
CountryBolivia

The Icla Formation is an early Devonian formation from Bolivia. It preserves a moderately diverse fauna of trilobites, brachiopods, crinoids and a few species of jawed fish.

Paleobiota

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
Paleobiota
Genus Species Higher taxon Notes Images
Ramirosuarezia R. boliviana Gnathostomata Resembles a mix of gnathostome groups, may be the same as Zamponiopteron[1]
Pucapampella P. rodrigae Stem-Chondrichthyes Also known from the Belèn Formation, one of the most basal known cartilaginous fish[2]
Fossilized jaws and teeth of P. rodrigae (A-C, G-H) and cartilage elements of Zamponiopteron (D-F), which may belong to the same animal, from the Department of Puno, Peru. Scale bar = 0.5 cm
Meperocrinus M. angelina Emperocrinidae (Camerata) Has unusual wing-shaped extensions of its calyx[3]
Lutocrinus L. boliviaensis Rhodocrinitidae (Diplobathrida)[4]
Griphocrinus G. pirovanoi Dimerocrinitidae (Diplobathrida)[4]
Maurotarion M. dereimsi, M. racheboufi Aulacopleuridae (Proetida)[5]
Bainella B. (Belenops) insolita Calmoniidae[6]
Deltacephalaspis D. (Prestalia) tumida Calmoniidae[6]
Schizostylus S. (Schizostylus) ?brevicaudatus Calmoniidae[6]
Probolops P. glabellirostris Calmoniidae[6]
Kozlowskiaspis K. (Kozlowskiapis) superna Calmoniidae[6]
Acastoides? ?A. verneuili Calmoniidae[6]
Phacopina P. padilla Calmoniidae[6]
Plesiomalvinella P. pujravii, P. boulei Calmoniidae[7]
Malvinella M. haugi Calmoniidae[7]
Vogesina V. devonica, V. lacunafera Calmoniidae Formerly classed in the genus Phacopina[7]
Nuculites N. beneckei, N. oblongatus[8] Malletiidae
Palaeoneilo P. rhysa?[8] Malletiidae
Phestia P. inornata?[8] Nuculanidae
Bolitheca B. steinmanni Orthothecida[9]
Bolithes B. crasquinae Hyolithida[10]
Protoleptostrophia P. concinna Stropheodontidae (Strophomenoidea)[11]
Schuchertella S. sp? Schuchertellidae (Orthotetida)[11]
Australostrophia A. mesembrina Chonostrophiidae (Chonetidina)[11]
Pleurochonetes P. anteloi, P. lauriata Chonetidae (Chonetidina)[11] P. anteloi was formerly placed in its own genus (Gamonetes) before being synonymised[12]
Notiochonetes N. falklandica Chonetidae (Chonetidina)[11]
Kentronetes K. iclaense[13] Chonetidina Formerly included in Quadrikentron, but split out into its own genus[14]
Antelocoelia A. johnsoni Leptocoeliidae (Rhynchonellida)[15]
Australocoelia A. palmata Leptocoeliidae (Rhynchonellida)[15]
Meristelloides M. rikowskii Meristellidae (Athyridida)[15]
Australospirifer A. hawkinsi Delthyrididae (Spiriferida)[15]
Metaplasia M. pseudoumbonata Ambocoeliidae (Spiriferida)[15]
Plicoplasia P. planoconvexa Ambocoeliidae (Spiriferida)[15]
Proboscidina P. arcei Centronellidae (Terebratulida)[15]
Mutationella M. sp Mutationellidae (Terebratulida)[15]
Podolella P. sp Mutationellidae (Terebratulida)[15]
Derbyina D. smithi, D. jamesiana Mutationellidae (Terebratulida)[15]
Pleurothyrella P. knodi Mutationellidae (Terebratulida)[15]
Scaphiocoelia S. boliviana Mutationellidae (Terebratulida)[15]
Cryptonella C. baini Cryptonellidae (Terebratulida)[15]
Conularia C. quichua, C. albertensis[16] Conulariida
Paraconularia P. ulrichiana, P. africana[16] Conulariida

References

  1. ^ Pradel, Alan; Maisey, John G.; Tafforeau, P.; Janvier, Philippe (May 2009). "An enigmatic gnathostome vertebrate skull from the Middle Devonian of Bolivia". Acta Zoologica. 90 (s1): 123–133. doi:10.1111/j.1463-6395.2008.00350.x.
  2. ^ Maisey, John G.; Janvier, Philippe; Pradel, Alan; Denton, John S. S.; Bronson, Allison; Miller, Randall; Burrow, Carole J. (31 December 2018). "Doliodus and Pucapampellids: Contrasting Perspectives on Stem Chondrichthyan Morphology". Evolution and Development of Fishes: 87–109. doi:10.1017/9781316832172.006.
  3. ^ Scheffler, Sandro M. (January 2021). "Crinoids from the Lower (Pragian–Emsian) and Middle (early Eifelian) Devonian of Bolivia (Icla and Belén formations, Malvinokaffric Realm)". Journal of Paleontology. 95 (1): 141–153. doi:10.1017/jpa.2020.70.
  4. ^ a b Thompson, Jeffrey R.; Ausich, William I.; Smith, Legrand (January 2013). "Echinoderms from the Lower Devonian (Emsian) of Bolivia (Malvinokaffric Realm)". Journal of Paleontology. 87 (1): 166–175. doi:10.1666/12-068R.1.
  5. ^ Adrain, Jonathan M.; Edgecombe, Gregory D. (1996). "Devonian Aulacopleurid trilobitesof the Malvinokaffric realm". Geobios. 29 (4): 417–436. doi:10.1016/S0016-6995(96)80003-9.
  6. ^ a b c d e f g Eldredge, Niles; Branisa, Leonardo (1980). "Calmoniid trilobites of the Lower Devonian Scaphiocoelia zone of Bolivia, with remarks on related species". Bulletin of the AMNH. 165 (2): 183–289.
  7. ^ a b c Lieberman, Bruce S.; Edgecombe, Gregory D.; Eldredge, Niles (1991). "Systematics and Biogeography of the "Malvinella Group," Calmoniidae (Trilobita, Devonian)" (PDF). Journal of Paleontology. 65 (5): 824–843. ISSN 0022-3360.
  8. ^ a b c Farjat, Alejandra (January 2018). SILURIAN AND DEVONIAN PROTOBRANCH BIVALVES FROM BOLIVIA (PDF). Asociación Boliviana de Paleontología. pp. 93–114.
  9. ^ Marek, L.; Isaacson, P. E. (1992). "Devonian Hyoliths from Bolivia". Journal of Paleontology. 66 (1): 24–28. ISSN 0022-3360.
  10. ^ Malinky, John M.; Racheboeuf, Patrick R. (2011). "New Hyolitha from the Devonian of Bolivia". Journal of Paleontology. 85 (6): 1077–1088. ISSN 0022-3360.
  11. ^ a b c d e Isaacson, Peter (January 1977). "Devonian stratigraphy and brachiopod paleontology of Bolivia. Part A. Orthida and Strophomenida". Palaeontographica Abteilung A. 155: 133–192.
  12. ^ Racheboeuf, Patrick R. (1998). The Chonetoidean Brachiopods. A revised and updated systematic and bibliographic catalogue. pp. 3–178.
  13. ^ Racheboeuf, Patrick R.; Branisa, Leonardo (1985). "New Data on Silurian and Devonian Chonetacean Brachiopods from Bolivia" (PDF). Journal of Paleontology. 59 (6): 1426–1450. ISSN 0022-3360.
  14. ^ Racheboeuf, Patrick R.; Herrera, Zarela (1 September 1994). "On some new Malvinokaffric Silurian and Devonian chonetacean brachiopods, and reclassification of others". Neues Jahrbuch für Geologie und Paläontologie - Monatshefte. 1994 (9): 541–560. doi:10.1127/njgpm/1994/1994/541.
  15. ^ a b c d e f g h i j k l m Isaacson, Peter (December 1993). "Devonian Brachiopoda of Bolivia". Fósiles y Facies de Bolivia. 13–14 (1–4): 3–55.
  16. ^ a b Babcock, Loren E.; Feldmann, Rodney M.; Wilson, Margaret T.; Suárez-Riglos, Mario (January 1987). "Devonian conulariids of Bolivia". National Geographic Research. 3 (2): 210–231.

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