Description: "Tegument covered by spines. Oral sucker not armed or armed by circumoral spines. Pharynx presented. Genital synus presented. Ventral and genital suckers usually not combined. Cirrus and bursa absent. Two testes located in posterior part of the body. Vitellaria in posterior part of the body."
^Leiper, R. T. (1909). London School of Tropical Medicine. "Report of the helminthologist for the six months ending 30th April, 1908". In, Report of the Advisory Committee for the Tropical Diseases Research Fund for the Year 1908 (pp. 35–39). London: Tropical Diseases Research Fund, Advisory Committee.