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Hemihegetotherium; Temporal range: Middle-Late Miocene ; ~12–7 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
	Skull of Hemihegetotherium achataleptum
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	†Notoungulata
Family:	†Hegetotheriidae
Subfamily:	†Hegetotheriinae
Genus:	†Hemihegetotherium; Rovereto 1914
Type species
	†Hemihegetotherium achataleptum; Rovereto, 1914
Species
	H. achataleptum Rovereto 1914; H. tantillum Vera 2019; H. torresi Cabrera & Kraglievich 2019;

Hemihegetotherium is an extinct genus of hegetotheriid notoungulate that lived from the Middle to the Late Miocene of what is now Argentina.

Description

It was a medium-sized animal and had long legs. Although not as those of modern lagomorphs, to which it was, in other regards, superficially similar. Like some of its relatives, the first pair of incisors of Hemihegetotherium was large, and it had simplified but high-crowned (hypsodont) molars and premolars. The structure of those teeth were so simplified in Hemihegetotherium that they consisted of simply a curved cylinder of enamel filled with dentin ; compared to Hegetotherium, the molars were more convex, with a semi-elliptical section. However, those teeth had pointed cusps along their outer margin. Hemihegetotherium had a tibia and fibula solidly fused together in their extremities, both distal and proximal.

Classification

The genus Hemihegetotherium was first described in 1914 by Gaetano Rovereto, based on fossil remains discovered in the Las Floras Formation (San Juan Province, Argentina) ; the type species is Hemihegetotherium achataleptum. The other species are H. torresi, from the Late Miocene of Argentina, and H. trilobus, from the Middle Miocene of Quebrada Honda in Bolivia, this last species distinguished from the other species of the genus by the presence of a trilobed third lower molar, hence its species name. H. tantillum, described in 2019, shared similarities with H. trilobus, and was found in Patagonia.

Hemihegetotherium was a member of Hegetotheriidae (hence its generic name, meaning "half Hegetotherium"), a group of usually small notoungulates with the superficial appearance of rodents or hares. Hemihegetotherium was similar but less specialized than the eponymous genus Hegetotherium.

Paleoecology

Hemihegetotherium was a terrestrial herbivore, probably consumming low and particularly fibrous plants, both in open and forested areas. The limbs of Hemihegetotherium may have been suitable both for running and digging.

Bibliography

Croft DA, Anaya F (2006) A new middle Miocene hegetotheriid (Notoungulata: Typotheria) and a phylogeny of the Hegetotheriidae. Journal of Vertebrate Paleontology 26:387–399.
F. D. Seoane, S. R. Juñent, and E. Cerdeño. 2017. Phylogeny and paleobiogeography of Hegetotheriidae (Mammalia, Notoungulata). Journal of Vertebrate Paleontology 37(1):e1278547:1-13
B. Vera (2019). "A new species and the record of Hemihegetotherium (Notoungulata, Hegetotheriidae) in the middle to Late Miocene of Patagonia, Argentina". Journal of South American Earth Sciences. 93: 23–35. doi:10.1016/j.jsames.2019.04.017.