^ abcVon Konrat, Matthew; Hentschel, Jörn; Heinrichs, Jochen; Braggins, John E.; Pócs, Tamás (1 November 2010). "Forty-one degrees below and sixty years in the dark: Frullania sect. Inconditum, a new section of Australasian Frullania species including F. colliculosa, sp. nov., F. hodgsoniae, nom. and stat. nov., F. aterrima, and F. hattorii (Frullaniaceae, Marchantiophyta)". Nova Hedwigia: 471–500. doi:10.1127/0029-5035/2010/0091-0471.