The Dysderoidea are a clade or superfamily of araneomorph spiders. The monophyly of the group, initially consisting of the four families Dysderidae, Oonopidae, Orsolobidae and Segestriidae,[2] has consistently been recovered in phylogenetic studies.[1][3] In 2014, a new family, Trogloraptoridae, was created for a recently discovered species Trogloraptor marchingtoni. It was suggested that Trogloraptoridae may be the most basal member of the Dysderoidea clade.[1][4] However, a later study found that Trogloraptoridae was placed outside the Dysderoidea and concluded that it was not part of this clade.[5]
Phylogeny
Dysderoidea are members of the Haplogynae clade: spiders with simpler copulatory organs (palpal bulbs and epigynes) than other araneomorphs. One hypothesis for relationships within the Haplogynae is shown below.[6] The status of the Trogloraptoridae is unclear. The family was not included in one study which otherwise found the same topography,[7] but it was placed outside even the Filistatidae in a 2014 study based on ribosomal DNA.[5]
Other studies have suggested that Caponiidae rather than Tetrablemmidae are the sister of Dysderoidea.[8]
References
^ abcMichalik, Peter & Ramírez, Martín J. (2014), "Evolutionary morphology of the male reproductive system, spermatozoa and seminal fluid of spiders (Araneae, Arachnida)–Current knowledge and future directions", Arthropod Structure & Development, 43 (4): 291–322, doi:10.1016/j.asd.2014.05.005, hdl:11336/19081, PMID24907603
^Forster, R.R. & Platnick, N.I. (1985), "A review of the austral spider family Orsolobidae (Arachnida, Araneae) with notes on the superfamily Dysderoidea", Bulletin of the American Museum of Natural History (181)
^Coddington, Jonathan A. & Levi, Herbert W. (1991), "Systematics and evolution of spiders (Araneae)", Annual Review of Ecology and Systematics, 22: 565–592, doi:10.1146/annurev.es.22.110191.003025, JSTOR2097274
^Griswold, C.; Audisio, T. & Ledford, J. (2012), "An extraordinary new family of spiders from caves in the Pacific Northwest (Araneae, Trogloraptoridae, new family)", ZooKeys (215): 77–102, doi:10.3897/zookeys.215.3547, PMC3428790, PMID22936872
^Coddington, Jonathan A. (2005), "Phylogeny and classification of spiders"(PDF), in Ubick, D.; Paquin, P.; Cushing, P.E. & Roth, V. (eds.), Spiders of North America: an identification manual, American Arachnological Society, pp. 18–24, retrieved 2015-09-24