The Dysderoidea are a clade or superfamily of araneomorph spiders. The monophyly of the group, initially consisting of the four families Dysderidae, Oonopidae, Orsolobidae and Segestriidae,[2] has consistently been recovered in phylogenetic studies.[1][3] In 2014, a new family, Trogloraptoridae, was created for a recently discovered species Trogloraptor marchingtoni. It was suggested that Trogloraptoridae may be the most basal member of the Dysderoidea clade.[1][4] However, a later study found that Trogloraptoridae was placed outside the Dysderoidea and concluded that it was not part of this clade.[5]
Dysderoidea are members of the Haplogynae clade: spiders with simpler copulatory organs (palpal bulbs and epigynes) than other araneomorphs. One hypothesis for relationships within the Haplogynae is shown below.[6] The status of the Trogloraptoridae is unclear. The family was not included in one study which otherwise found the same topography,[7] but it was placed outside even the Filistatidae in a 2014 study based on ribosomal DNA.[5]
Trogloraptoridae
Filistatidae
Caponiidae
Tetrablemmidae
Segestriidae
Dysderidae
Orsolobidae
Oonopidae
remaining haplogynes
Other studies have suggested that Caponiidae rather than Tetrablemmidae are the sister of Dysderoidea.[8]
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