Pine forests are found primarily in well-drained, nutrient-poor, acidic soils such as quartziferoussands, pseudo-spodosols in the west, and lateritic soils. Pine trees and encino (Quercus sagrana) obtain nutrients through an ectomycorrhizal symbiosis with fungi, allowing them to attain tree size. The forests feature a dense xerophytic brushy story of mainly Rubiaceae, Euphorbiaceae, Myrtaceae, and Melastomataceae along with a herbaceous story of a few epiphytes, primarily from the genus Tillandsia, and lianas. Secondary forests formed by deforestation have a more open canopy with an understory dominated by Comocladia dentata; grasses, lianas and epiphytes are poorly represented.[2]
Small patches of pine forest occur around the Sierra Cristal, Nipe-Sagua-Baracoa Mountains, and Sierra Maestra. These forests are found at higher elevations than western pine forests, up to 800–1,800 m (2,600–5,900 ft), making them cooler and wetter. Consequently, some of the vegetation resembles that of the rainforests, with an abundance of ferns. Pino de la Maestra (Pinus maestrensis) predominates in the Sierra Maestra, developing on landslide areas over granitic rock, while the Pino de Mayarí (Pinus cubensis) predominates in the Nipe-Sagua-Baracoa. Some accompanying species in these forests are the griñapo (Dracaena cubensis), Eupatorium spp., Myrtus spp., Baccharis spp., Jacaranda arborea and Eugenia pinetorum.[2]
^World Wildlife Fund, ed. (2001). "Cuban pine forests". WildWorld Ecoregion Profile. National Geographic Society. Archived from the original on 2010-03-08. Retrieved 2010-11-21.