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Copiula
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Amphibia
Order:	Anura
Family:	Microhylidae
Subfamily:	Asterophryinae
Genus:	Copiula; Méhelÿ, 1901
Type species
	Phrynixalus oxyrhinus; Boulenger, 1898
Species
	14 species (see text)

Copiula is a genus of microhylid frogs endemic to New Guinea. The common name Mehely frogs has been coined for them.^[1] They are leaf-litter inhabitants.^[2]

Taxonomy

Copiula is probably not monophyletic. Some former Austrochaperina species have already been transferred to this genus, and further ones might follow when more data became available.^[1]^[3]

Species

There are at present 14 species in this genus:^[1]

Copiula alpestris (Zweifel, 2000)
Copiula annanoreenae Günther, Richards, and Dahl, 2014
Copiula derongo (Zweifel, 2000)
Copiula exspectata Günther, 2002
Copiula fistulans Menzies and Tyler, 1977 — Lae Mehely Frog
Copiula guttata (Zweifel, 2000)
Copiula lennarti Günther, Richards, and Dahl, 2014
Copiula major Günther, 2002
Copiula minor Menzies and Tyler, 1977 — Milne Bay Mehely Frog
Copiula obsti Günther, 2002
Copiula oxyrhina (Boulenger, 1898) — Misima Island Mehely Frog
Copiula pipiens Burton and Stocks, 1986 — Wirui Mehely Frog
Copiula rivularis (Zweifel, 2000)
Copiula tyleri Burton, 1990 — Sepik Mehely Frog

The AmphibiaWeb^[4] reports fewer species, with species that Peloso and colleagues moved in 2016 from Austrochaperina and Oxydactyla missing.^[3]

References

^ ^a ^b ^c Frost, Darrel R. (2016). "Copiula Méhely, 1901". Amphibian Species of the World: an Online Reference. Version 6.0. American Museum of Natural History. Retrieved 20 July 2016.
^ Zweifel, R. G. (2000). "Partition of the Australopapuan microhylid frog genus Sphenophryne with descriptions of new species". Bulletin of the American Museum of Natural History. 253: 1–130. doi:10.1206/0003-0090(2000)253<0001:POTAMF>2.0.CO;2. hdl:2246/1600. S2CID 85621508.
^ ^a ^b Peloso, Pedro L.V.; Frost, Darrel R.; Richards, Stephen J.; Rodrigues, Miguel T.; Donnellan, Stephen; Matsui, Masafumi; Raxworthy, Cristopher J.; Biju, S.D.; Lemmon, Emily Moriarty; Lemmon, Alan R.; Wheeler, Ward C. (2016). "The impact of anchored phylogenomics and taxon sampling on phylogenetic inference in narrow-mouthed frogs (Anura, Microhylidae)". Cladistics. 32 (2): 113–140. doi:10.1111/cla.12118. PMID 34732021. S2CID 84925667.
^ "Microhylidae". AmphibiaWeb: Information on amphibian biology and conservation. [web application]. Berkeley, California: AmphibiaWeb. 2016. Retrieved 20 July 2016.

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[Frost-1] Frost, Darrel R. (2016). "Copiula Méhely, 1901". Amphibian Species of the World: an Online Reference. Version 6.0. American Museum of Natural History. Retrieved 20 July 2016.

[Zweifel-2] Zweifel, R. G. (2000). "Partition of the Australopapuan microhylid frog genus Sphenophryne with descriptions of new species". Bulletin of the American Museum of Natural History. 253: 1–130. doi:10.1206/0003-0090(2000)253<0001:POTAMF>2.0.CO;2. hdl:2246/1600. S2CID 85621508.

[Peloso_et_al._2016-3] Peloso, Pedro L.V.; Frost, Darrel R.; Richards, Stephen J.; Rodrigues, Miguel T.; Donnellan, Stephen; Matsui, Masafumi; Raxworthy, Cristopher J.; Biju, S.D.; Lemmon, Emily Moriarty; Lemmon, Alan R.; Wheeler, Ward C. (2016). "The impact of anchored phylogenomics and taxon sampling on phylogenetic inference in narrow-mouthed frogs (Anura, Microhylidae)". Cladistics. 32 (2): 113–140. doi:10.1111/cla.12118. PMID 34732021. S2CID 84925667.

[AmphibiaWeb-4] "Microhylidae". AmphibiaWeb: Information on amphibian biology and conservation. [web application]. Berkeley, California: AmphibiaWeb. 2016. Retrieved 20 July 2016.

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