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Archipolypoda; Temporal range: Lochkovian–Pennsylvanian PreꞒ Ꞓ O S D C P T J K Pg N
	Reconstruction of Acantherpestes major
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Subphylum:	Myriapoda
Class:	Diplopoda
Subclass:	Chilognatha
Superorder:	†Archipolypoda; Scudder, 1882
Orders
	Archidesmida; Cowiedesmida; Euphoberiida; Palaeosomatida
Synonyms
	Macrosterni Fritsch, 1899; Palaeocoxopleura Verhoeff, 1928

Archipolypoda is an extinct group of millipedes known from fossils in Europe and North America and containing the earliest known land animals.^[1] The Archipolypoda was erected by Scudder (1882)^[2] but redefined in 2005 with the description of several new species from Scotland.^[3] Distinguishing characteristics include relatively large eyes with densely packed ocelli (sometimes interpreted as compound eyes), and modified leg pairs on the 8th body ring.^[3]^[4] Some species had prominent spines while others had a flattened appearance.^[3]^[5]

Classification

The Archipolypoda as currently recognized consists of four orders,^[6] many with monotypic families and genera, as well as five species of uncertain placement (incertae sedis).^[3]^[7]

Archidesmida Wilson & Anderson, 2004

Archidesmidae Scudder, 1885
- Archidesmus macnicoli Peach, 1882. Lower Devonian, Scotland
Zanclodesmidae Wilson, Daeschler & Desbiens, 2005
- Zanclodesmus willetti Wilson, Daeschler & Desbiens, 2005. Upper Devonian, Quebec, Canada
- Orsadesmus rubecollus Wilson, Daeschler & Desbiens, 2005. Upper Devonian, Pennsylvania, USA

Cowiedesmida Wilson & Anderson, 2004

Cowiedesmidae Wilson & Anderson, 2004
- Cowiedesmus eroticopodus Wilson & Anderson, 2004. Mid Silurian or Lower Devonian,^[8] Scotland.

Euphoberiida Hoffman, 1969

Euphoberiidae Scudder, 1882. Upper Carboniferous of Europe and North America.
- Acantherpestes Meek & Worthen, 1868
- Euphoberia Meek & Worthen, 1868
- Myriacantherpestes Burke, 1979

Palaeosomatida Hannibal & Krzeminski, 2005. Carboniferous, UK and Poland^[9]

Palaeosomatidae Hannibal & Krzeminski, 2005
- Palaeosoma

Order incertae sedis

Albadesmus almondi Wilson & Anderson, 2004 Mid Silurian or Lower Devonian,^[8] Scotland
Anaxeodesmus diambonotus Wilson, 2005 Upper Carboniferous, UK.
Anthracodesmus macconochiei, Peach, 1899
Palaeodesmus tuberculata (Brade-Birks, 1923) (=Kampecaris tuberculata) Lower Devonian, Scotland.
Pneumodesmus newmani Wilson & Anderson, 2004 Mid Silurian or Lower Devonian,^[8] Scotland

References

^ Selden, Paul; Helen Read (2008). "The Oldest Land Animals: Silurian Millipedes from Scotland" (PDF). Bulletin of the British Myriapod & Isopod Group. 23: 36–37.
^ Scudder, Samuel H. (1882). "Archipolypoda, a Subordinal Type of Spined Myriapods from the Carboniferous Formation". Memoirs of the Boston Society of Natural History. 3 (5): 143.
^ ^a ^b ^c ^d Wilson, Heather M.; Anderson, Lyall I. (2004). "Morphology and taxonomy of Paleozoic millipedes (Diplopoda: Chilognatha: Archipolypoda) from Scotland". Journal of Paleontology. 78 (1): 169–184. Bibcode:2004JPal...78..169W. doi:10.1666/0022-3360(2004)078<0169:MATOPM>2.0.CO;2. JSTOR 4094847. S2CID 131201588.
^ Sierwald, Petra; Bond, Jason E. (2007). "Current Status of the Myriapod Class Diplopoda (Millipedes): Taxonomic Diversity and Phylogeny". Annual Review of Entomology. 52 (1): 401–420. doi:10.1146/annurev.ento.52.111805.090210. PMID 17163800.
^ Wilson, Heather M.; Daeschler, Edward B.; Desbiens, Sylvain (2005). "New Flat-Backed Archipolypodan Millipedes from the Upper Devonian of North America". Journal of Paleontology. 79 (4): 738–744. doi:10.1666/0022-3360(2005)079[0738:NFAMFT]2.0.CO;2. JSTOR 4095046. S2CID 140190612.
^ Shear, William A.; Edgecombe, Gregory D. (2010). "The geological record and phylogeny of the Myriapoda". Arthropod Structure & Development. 39 (2–3): 174–190. Bibcode:2010ArtSD..39..174S. doi:10.1016/j.asd.2009.11.002. PMID 19944188.
^ Wilson, Heather M. (2005). "A new genus of Archipolypodan millipede from the Coseley Lagerstatte, Upper Carboniferous, UK". Palaeontology. 48 (5): 1097–1100. Bibcode:2005Palgy..48.1097W. doi:10.1111/j.1475-4983.2005.00496.x.
^ ^a ^b ^c Suarez, Stephanie E.; Brookfield, Michael E.; Catlos, Elizabeth J.; Stöckli, Daniel F. (2017-06-28). "A U-Pb zircon age constraint on the oldest-recorded air-breathing land animal". PLOS ONE. 12 (6): e0179262. Bibcode:2017PLoSO..1279262S. doi:10.1371/journal.pone.0179262. ISSN 1932-6203. PMC 5489152. PMID 28658320.
^ Hannibal, Joseph T.; Krzeminski, Wieslaw (2005). "A palaeosomatid millipede (Archipolypoda: palaeosomatida) from the Carboniferous (Namurian A) of Silesia, Poland". Polskie Pismo Entomologiczne. 74 (3): 205–217.