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Anhanguerids; Temporal range: Early-Late Cretaceous, 140–92.5 Ma PreꞒ Ꞓ O S D C P T J K Pg N
	Anhanguera santanae skeleton
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Order:	†Pterosauria
Suborder:	†Pterodactyloidea
Clade:	†Anhangueria
Family:	†Anhangueridae; Campos & Kellner 1985
Type species
	†Anhanguera blittersdorffi; Campos & Kellner, 1985
Subgroups
	†Anhanguerinae †Anhanguera; †Araripesaurus?; †Arthurdactylus?; †Brasileodactylus?; †Caulkicephalus?; †Cearadactylus?; †Guidraco?; †Liaoningopterus; †Ludodactylus?; †Maaradactylus; †Santanadactylus?; ; †Coloborhynchinae? †Aerodraco?; †Coloborhynchus; †Nicorhynchus; †Ornithocheirus?; †Siroccopteryx?; †Uktenadactylus; ; Tropeognathinae?;

Anhangueridae is a group of pterosaurs within the suborder Pterodactyloidea.^[2] They were among the last pterosaurs to possess teeth. A recent study discussing the group considered the Anhangueridae to be typified by a premaxillary crest and a lateral expansion in the distal rostrum. The same study presented a cladistic analysis, for which an "agreement subtree" was calculated. The Anhangueridae was found to be sister taxon to the large crested Tropeognathus.^[3]

Relationships

There are competing theories of ornithocheiromorph phylogeny (evolutionary relationships). Below is cladogram following a topology recovered by Brian Andres, using the most recent iteration of his data set.^[4]

Anhangueridae

Aerodraco sedgwickii

Liaoningopterus gui

Anhanguera

Anhanguera robustus

	Anhanguera piscator

	Anhanguera blittersdorffi

The cladogram below follows Pêgas et al. (2019), who recovered Anhangueridae as a much more inclusive group. The analysis found most of the ornithocheirids falling into this family, while Ornithocheirus itself was recovered as a basal member of Ornithocheirae.^[5]

Anhangueria

Hamipteridae

	Hamipterus

	Iberodactylus

Anhangueridae

Tropeognathus

Coloborhynchinae

	Coloborhynchus

	Siroccopteryx

	Uktenadactylus

Anhanguerinae

	Caulkicephalus

	Guidraco

	Ludodactylus

Anhanguera

Liaoningopterus

	Cearadactylus

	Maaradactylus

The clagogram below is reproduced from Richards et al., (2023), who based their data matrix on the data matrix in Holgado and Pêgas, (2020). In the paper they erect the clade Mythungini which comprises all Australian tropeognathines.^[6]