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Thursophyton; Temporal range: Middle Devonian PreꞒ Ꞓ O S D C P T J K Pg N
	Axes of Thursophyton milleri, detail from Nathorst (1915), Fig. 7
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Order:	incertae sedis
Genus:	†Thursophyton; Nath. 1915
Type species
	Thursophyton milleri; (J.W.Salter) Nath.

Thursophyton ("plant from Thurso") is a form genus of extinct vascular plants known from anatomically preserved specimens originally described from the Givetian (Middle Devonian) of Scotland, and since reported from other parts of northern Europe.^[2] The taxonomic position of Thursophyton has been uncertain; most authors have considered it too imperfectly known to place, or have assigned it to the Lycopodiophyta. Unpublished research suggests similarities with spiny genera of the Zosterophyllopsida.^[3]

Description

Thursophyton is a genus of terrestrial vascular plants which flourished in the Middle Devonian period. These plants consisted of aerial stems branching dichotomously, trichotomously or pseudomonopodially, at least the main axes clothed in spirally arranged spines up to 7 mm long, which are not leaves as they are not vascularised and leave no scar when removed.^[2] The stems contain an elliptical exarch xylem having both annular and spirally thickened tracheids; very little is known about the sporangia.^[3]

Species

The type species is Thursophyton milleri. Other species have been suggested (e.g. T. reidii, T. eberfeldense, T. hostimense, T. vahlbergianum) which are usually regarded as synonyms of T. milleri. Because of the poor preservation of much of this plant material from the Devonian period, it is likely that other synonyms exist e.g. Euthursophyton hamperbachense Mustafa from Germany.^[3]