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Stipeae
	Celtica gigantea
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Clade:	Angiosperms
Clade:	Monocots
Clade:	Commelinids
Order:	Poales
Family:	Poaceae
Clade:	BOP clade
Subfamily:	Pooideae
Supertribe:	Stipodae
Tribe:	Stipeae; Dumort.
Genera
	28 genera, see text
Synonyms
	supertribe Stipodae L. Liu; subtribe Stipinae Griseb.; Aciachninae Caro; Ortachninae Caro;

The Stipeae are a tribe of grasses within the subfamily Pooidae, with up to 600 described species.^[2]

Description

The defining morphological features of the Stipeae include single-flowered spikelets lacking a rachilla extension, and the lemmas (the external bract) have either a sharp point or a terminal awn (long bristle).^[3]

Genera

The tribe includes 32 genera:^[1]^[4]^[5]

Achnatherum P.Beauv. (syn. Aristella)
Aciachne Benth.
Amelichloa Arriaga & Barkworth
Anemanthele Veldkamp
Austrostipa S.W.L.Jacobs & J.Everett
Barkworthia Romasch., P.M.Peterson & Soreng
Celtica F.M.Vázquez & Barkworth
Eriocoma Nutt.
× Eriosella Romasch.
Hesperostipa (Elias) Barkworth
Jarava Ruiz & Pav.
Lorenzochloa Reeder & C.Reeder
Macrochloa Kunth
Nassella (Trin.) É.Desv.
Neotrinia (Tzvelev) M.Nobis, P.D.Gudkova & A.Nowak
Oloptum Röser & Hamasha
Ortachne Nees ex Steud.
Orthoraphium Nees
Oryzopsis Michx.
Pappostipa (Speg.) Romasch., P.M.Peterson & Soreng
Patis Ohwi
Piptatheropsis Romasch., P.M.Peterson & Soreng
Piptatherum P.Beauv.
Piptochaetium J.Presl.
Psammochloa Htchc.
Pseudoeriocoma Romasch., P.M.Peterson & Soreng
Ptilagrostiella Romasch., P.M.Peterson & Soreng
Ptilagrostis Griseb.
Stipa L.
Stipellula Röser & Hamasha (syn. Stipella)
Thorneochloa Romasch., P.M.Peterson & Soreng
Timouria Roshev.
Trikerala Bor

Many species initially placed into Stipa have now been split off into new genera. Some recent papers have analysed relationships within and between the genera,^[6]^[7]^[8] but a complete analysis has not yet been performed. Stipoid grasses use the C₃ photosynthetic pathway and live in temperate areas worldwide.^[9]

Known fossils date from the late Miocene.^[10]

References

^ ^a ^b Soreng, Robert J.; Peterson, Paul M.; Romaschenko, Konstantin; Davidse, Gerrit; Teisher, Jordan K.; Clark, Lynn G.; Barberá, Patricia; Gillespie, Lynn J.; Zuloaga, Fernando O. (2017). "A worldwide phylogenetic classification of the Poaceae (Gramineae) II: An update and a comparison of two 2015 classifications". Journal of Systematics and Evolution. 55 (4): 259–290. doi:10.1111/jse.12262. hdl:10261/240149. ISSN 1674-4918.
^ Barkworth ME, Arriaga MO, Smith JF, Jacobs SWL, Valdes-Reyna J, Bushman BS (2008) Molecules and Morphology in South American Stipeae (Poaceae). Systematic Botany 33(4), 719-731.
^ Cialdella AM, Giussani LM, Aagesen L, Zuloaga FO, Morrone O (2007) A phylogeny of Piptochaetium (Poaceae : Pooideae : Stipeae) and related genera based on a combined analysis including trnL-F, rp116, and morphology. Systematic Botany 32(3), 545-559.
^ Peterson PM, Romaschenko K, Soreng RJ, Valdés Reyna J (2019) A key to the North American genera of Stipeae (Poaceae, Pooideae) with descriptions and taxonomic names for species of Eriocoma, Neotrinia, Oloptum, and five new genera: Barkworthia, × Eriosella, Pseudoeriocoma, Ptilagrostiella, and Thorneochloa. PhytoKeys 126: 89-125. https://doi.org/10.3897/phytokeys.126.34096
^ "Poaceae Barnhart". Plants of the World Online. Royal Botanic Gardens, Kew. Retrieved 27 October 2024.
^ Barber, Janet C.; Hames, Kasey A.; Cialdella, Ana Maria; Giussani, Liliana M.; Morrone, Osvaldo (2009). "Phylogenetic relationships of Piptochaetium Presl (Poaceae: Stipeae) and related genera reconstructed from nuclear and chloroplast sequence datasets". Taxon. 58 (2): 375–380. doi:10.1002/tax.582005. ISSN 0040-0262. pdf
^ Jacobs SWL, Bayer R, Everett J, Arriaga MO, Barkworth ME, Sabin-Badereau A, Torres MA, Vazquez FM, Bagnall N (2007) Systematics of the tribe Stipeae (Gramineae) using molecular data. Aliso 23, 349-361.
^ Jacobs SWL, Everett J, Barkworth ME, Hsiao C (2000) Relationships within the Stipoid grasses (Gramineae). In 'Grass Systematics and Evolution.' (Eds SWL Jacobs and J Everett) pp. 75-82. (CSIRO: Melbourne)
^ Romaschenko K, Peterson PM, Soreng RJ, Garcia-Jacas N, Futorna O, Susanna A (2008). "Molecular phylogenetic analysis of the American Stipeae (Poaceae) resolves Jarava sensu lato polyphyletic: evidence for a new genus, Pappostipa". Journal of the Botanical Research Institute of Texas. 2 (1): 165–192. JSTOR 41971613.{{cite journal}}: CS1 maint: multiple names: authors list (link) pdf
^ Thomasson, Joseph R. (2005). "Berriochloa Gabeli and Berrichloa Huletti (Graminae: Stipeae), two new grass species from the late miocene Ash Hollow Formationof Nebraska and Kansas". Journal of Paleontology. 79 (1): 185–199. Bibcode:2005JPal...79..185T. doi:10.1666/0022-3360(2005)079<0185:BGABHG>2.0.CO;2. ISSN 0022-3360. pdf