^ abEschmeyer, W. N.; R. Fricke; and R. van der Laan (7 April 2017). Catalog of Fishes. Retrieved 7 April 2017.
^ abVan Steenberge; Pariselle; Huyse; Volckaert; Snoeks; and Vanhove (2015). Morphology, Molecules, and Monogenean Parasites: An Example of an Integrative Approach to Cichlid Biodiversity. PLOS ONE 10(6): e0129987. doi:10.1371/journal.pone.0124474
^ abKonings, A.F. (2015). Tanganyika Cichlids in their natural habitat. 3rd edition. ISBN978-1932892185.